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Dioscorea pentaphylla

L. (1753)

This name is accepted

Kingdom: Viridiplantae Phylum: Magnoliophyta Class/Clade: Monocots Order: Dioscoreales Family: Dioscoreaceae Genus: Dioscorea

fiveleaf yam, pi‘a [pia] Hawai‘i

Description

Key Characters:

Growth Form: Vines.

Stems: Stems twining to the left, = angled, up to 10 m long, pilose to hirsute, glabrate with age, usually with numerous scattered prickles, tubers nearly always solitary, variable in shape, growing near the surface or over 1 m underground.

Roots: Fibrous root system.

Leaves: Leaves compound (palmate) or rarely simple. Alternate. Leaflets 3–5, elliptic or oblong, 5–10 cm long, 2–4 cm wide, the central one oblanceolate or obovate, the lateral ones somewhat asymmetrical. Apex obtuse or acute to acuminate. Surfaces with yellowish brown pubescent at least on lower surface, axils with bulbils and occasionally bearing globose to horseshoe shaped tubers ca. 1 cm in diameter. Leaflet margins entire. Petioles 5-11 cm long, densely reddish brown pubescent. Stipules absent.

Flowers: Staminate flowers in axillary, spike-like racemes up to 30 cm long, near the ends of the branches, reddish pubescent, pistillate flowers in long pendent spikes 5–25 cm long. Flowers unisexual, the plants dioecious. Calyx of staminate flowers with Tepals broadly lanceolate, ca. 1 mm long. Calyx of pistillate flowers with tepals ca. 1.5 mm long. Staminate flower with 3 stamen and 3 staminodes (longer than stamens); pistillate flowers with 6 rudimentary staminodes. Ovary inferior, villous, 3-carpellate.

Fruit: Capsules oblong; 20–25 mm long; glabrous or glabrate; shallowly emarginate at both ends; the wings up to 6 mm wide. Seeds winged on basal side only.

Ploidy: 2n = 40; 70; 80; 144

Habitat: Naturalized primarily in windward mesic valleys.

Elevation Range: 115–325 m.

Historical Distribution

Uses and Culture

USES

  • Tubers eaten in times of famine (Krauss 1993:11,12).

Natural History

Statewide Status

Naturalized

Island Status

Kaua'i Naturalized
O'ahu Naturalized
Molokai Naturalized
Maui Naturalized
Hawai'i Naturalized

Dispersal Agents


Pollinators

Specimens

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Bibliography

Name Published In: Sp. Pl.: 1032 (1753)

Other References

Wagner et al. 1990:1437 (K, O, Mo, M, H)

Occurrences

SNo. Scientific Name Scientific Name Authorship Locality Habitat Basis of Record Recorded By Record Number Island Source Date
1 Dioscorea pentaphylla L. Kuilau Trail, below Keahua Arboretum. Disturbed lowland windward Forest PRESERVED_SPECIMEN Natalia Tangalin 3559 Kauai PTBG 11/22/2013
2 Dioscorea pentaphylla L. Slopes of Waiohookalo Stream Valley (N. Molokai); steep, rocky slopes. Alt. 100-180 m. With secondary forest. PRESERVED_SPECIMEN David H. Lorence 5419 Molokai PTBG 7/31/1987
3 Dioscorea pentaphylla L. waimalu, koolay range, oahu, territory of hawaii PRESERVED_SPECIMEN o. degener 10110 Hawaii US 11/3/1935
4 Dioscorea pentaphylla L. coast nw of hilo PRESERVED_SPECIMEN wilkes explor. exped. s.n. Hawaii US
5 Dioscorea pentaphylla L. Waimalu, Koolau Range, at middle elevation PRESERVED_SPECIMEN Degener, O.; Park; Potter; Shigeura; Takamoto, -. 10110 Oahu US 11/3/1935
6 Dioscorea pentaphylla L. honaunau PRESERVED_SPECIMEN a. s. hitchcock 14545 Hawaii US 9/3/1916
7 Dioscorea pentaphylla L. waimalu, koolau range, at middle elevation PRESERVED_SPECIMEN o. degener, park, potter, shigeura & -. takamoto 10110 Oahu US 11/3/1935
8 Dioscorea pentaphylla L. Honaunau PRESERVED_SPECIMEN Hitchcock, A. S. 14545 Hawaii US 9/3/2016
9 Dioscorea pentaphylla L. 3.2 mi NW of Wailua Experiment Station, Lihue-Koloa Forest Reserve PRESERVED_SPECIMEN Ishikawa, S. S. 277 Kauai US 8/20/1973
10 Dioscorea pentaphylla L. oahu PRESERVED_SPECIMEN st.john, h. collector number: 24834 Oahu BISH
11 Dioscorea pentaphylla L. PRESERVED_SPECIMEN degener, otto 10110 USCH 11/3/1935
12 Dioscorea pentaphylla L. Wailau Valley Climbing over shrubs. PRESERVED_SPECIMEN Rock, J.F.C. 70221 Molokai BISH 4/20/1910
13 Dioscorea pentaphylla L. waioli valley PRESERVED_SPECIMEN forbes, c.n. collector number: 91.k Kauai BISH 7/29/1909
14 Dioscorea pentaphylla L. palolo valley PRESERVED_SPECIMEN forbes, c.n. collector number: 1968.o Oahu BISH 10/22/1914
15 Dioscorea pentaphylla L. East Maui, Hana District, Kalahu Point, PTBG Kahanu Garden In remnant Pandanus forest SSE of Pi`ilanihale Heiau in rough a`a lava. Common throughout forest. PRESERVED_SPECIMEN Flynn, T. 2487 Maui BISH 10/20/1987
16 Dioscorea pentaphylla L. koolau range; waimalu Flowering only bare landslide at middle elevation. PRESERVED_SPECIMEN degener, o. park, k.k.; potter; shigeura; takamoto collector number: 10110 Oahu BISH 11/3/1935
17 Dioscorea pentaphylla L. 3.2 mi nw of wailua experiment station, lihue-koloa forest reserve PRESERVED_SPECIMEN s. s. ishikawa 277 Kauai US 8/20/1973
18 Dioscorea pentaphylla L. head of hanakoa valley Wet woods. PRESERVED_SPECIMEN st.john, h. collector number: 24968 Kauai BISH 1/3/1953
19 Dioscorea pentaphylla L. Koolau Range; Palolo, Waiomao Valley Intermediate moist. PRESERVED_SPECIMEN Nishida, T. s.n. Oahu BISH 10/5/1943
20 Dioscorea pentaphylla L. punaluu; castle trail Loose rocky soil, steep slope, ohia woods. PRESERVED_SPECIMEN st.john, h. collector number: 23376 Oahu BISH 5/23/1948
21 Dioscorea pentaphylla L. Wailau Valley floor PRESERVED_SPECIMEN Higashino, P.K. 10374 Molokai BISH 8/13/1984
22 Dioscorea pentaphylla L. wainiha; wainiha valley, bottom of small side gulch Moist lower forest. Climbing over trees. PRESERVED_SPECIMEN st.john, h. collector number: 13908 Kauai BISH 1/1/1934
23 Dioscorea pentaphylla L. ewa dist.; waimalu Metrosideros forest. PRESERVED_SPECIMEN grant, m.l. collector number: 7338 Oahu BISH 9/9/1934
24 Dioscorea pentaphylla L. Waimano Trail PRESERVED_SPECIMEN Kerr, M. 439-1 Oahu BISH 10/7/1945
25 Dioscorea pentaphylla L. Koolau Range; Palolo, Waiomao Valley Climbing over low shrubs, dense undergrowth. PRESERVED_SPECIMEN Kuykendall, J.R. 45 Oahu BISH 11/29/1942
26 Dioscorea pentaphylla L. Gulch west of Ualapue Rocky woods. PRESERVED_SPECIMEN Degener, O. 5562 Molokai BISH 6/30/1928
27 Dioscorea pentaphylla L. east maui, hana district, 2.5 km w of hana hwy & waikoloa rd in the kawaipapa area Tropical lowland forest planted w. introduced trees. PRESERVED_SPECIMEN annable, c.r. hobdy, r. collector number: 3237 Maui BISH 1/15/1997
28 Dioscorea pentaphylla L. waimano trail PRESERVED_SPECIMEN kerr, m. collector number: 439-1 Oahu BISH 10/7/1945
29 Dioscorea pentaphylla L. waimalu Damp shaded rocks in moderately dry mesic ohia forest w/ Aleurites and Eugenia malaccensis common in wetter gullies PRESERVED_SPECIMEN brown, f.b.h. collector number: 11282 Oahu BISH 12/31/1925
30 Dioscorea pentaphylla L. ahupuaa of halepuaa forest reserve in the puna district. found growing on the side of the old railroad On the side of the old railroad. PRESERVED_SPECIMEN esp field crew collector number: s.n. Hawaii BISH 8/15/1979
31 Dioscorea pentaphylla L. ahupuaa of Halepuaa Forest Reserve in the Puna District. Found growing on the side of the old railroad On the side of the old railroad. PRESERVED_SPECIMEN ESP Field Crew s.n. Hawaii BISH 8/15/1979
32 Dioscorea pentaphylla L. head of wailau valley Rain forest near banana patch. PRESERVED_SPECIMEN st.john, h. et al. collector number: 23531 Molokai BISH 12/27/1948
33 Dioscorea pentaphylla L. Wainiha; Wainiha Valley, bottom of small side gulch Moist lower forest. Climbing over trees. PRESERVED_SPECIMEN St. John, H. 13908 Kauai BISH 1/1/1934
34 Dioscorea pentaphylla L. Palolo, Waiomao Valley Open shrubby area in valley. PRESERVED_SPECIMEN Tongg, R. s.n. Oahu BISH 11/29/1942
35 Dioscorea pentaphylla L. north molokai; slopes of waiahookalo stream valley Steep, rocky slopes w/secondary forest of Psidium guajava, Syzygium malaccense & native remnants incl. Diospyros sandwicensis, Metrosideros & Nestegis w/understory of Kalanchoe pinnata & Zingiber zerumbet. Common along stream. PRESERVED_SPECIMEN lorence, d.h. wagner, w.l.; perlman, s.; montgomery, s. collector number: 5419 Molokai BISH 7/31/1987
36 Dioscorea pentaphylla L. Head of Wailau Valley Rain forest near banana patch. PRESERVED_SPECIMEN St. John, H. 23531 Molokai BISH 12/27/1948
37 Dioscorea pentaphylla L. koolau range; palolo, waiomao valley Intermediate moist. PRESERVED_SPECIMEN nishida, t. collector number: s.n. Oahu BISH 10/5/1943
38 Dioscorea pentaphylla L. kapakahi gulch Koa zone in ope kukui forest. PRESERVED_SPECIMEN egler, f.e. collector number: 37-215 Oahu BISH 7/22/1937
39 Dioscorea pentaphylla L. lihue-koloa forest reserve; 3.2 miles nw of wailua experiment station Vine climbing on Aleurites & Hibiscus overstory. In shade w/Zingiber, Nephrolepis, Oplismenus. PRESERVED_SPECIMEN ishikawa, s. collector number: 277 Kauai BISH 8/20/1973
40 Dioscorea pentaphylla L. east maui, hana district, hana forest reserve, hana ranch lands, mauka of keakaamanu Alien forest slopes dominated by Psidium guajava and Ardisia elliptica. PRESERVED_SPECIMEN imada, c.t. legrande, m.; jansen, s.; crummer, j. collector number: 2002-80 Maui BISH 11/11/2002
41 Dioscorea pentaphylla L. Waimalu PRESERVED_SPECIMEN Brown, F.B.H. 11282 Oahu BISH 12/31/1925
42 Dioscorea pentaphylla L. Waioli Valley PRESERVED_SPECIMEN Forbes, C.N. 91.K Kauai BISH 7/29/1909
43 Dioscorea pentaphylla L. ahupuaa of Halepuaa Forest Reserve in the Puna District. Found growing on the side of the old railroad Tropical lowland forest, planted with introduced trees PRESERVED_SPECIMEN C. R. Annable 3237 Hawaii HAPI 1/15/1997
44 Dioscorea pentaphylla L. koolau range; palolo, waiomao valley Climbing over low shrubs, dense undergrowth. PRESERVED_SPECIMEN kuykendall, j.r. collector number: 45 Oahu BISH 11/29/1942
45 Dioscorea pentaphylla L. Koolau Range; Waimalu Flowering only bare landslide at middle elevation. PRESERVED_SPECIMEN Degener, O. 10110 Oahu BISH 11/3/1935
46 Dioscorea pentaphylla L. East Maui, Hana District, Hana Forest Reserve, Hana Ranch lands, mauka of Keakaamanu Tropical lowland forest, planted with introduced trees UNKNOWN C. R. Annable Maui HAPI 1/15/1997
47 Dioscorea pentaphylla L. Head of Hanakoa Valley Wet woods. PRESERVED_SPECIMEN St. John, H. 24968 Kauai BISH 1/3/1953
48 Dioscorea pentaphylla L. East Maui, Hana District, 2.5 km W of Hana Hwy & Waikoloa Rd in the Kawaipapa area Tropical lowland forest planted w. introduced trees. PRESERVED_SPECIMEN Annable, C.R. Hobdy, R. Collector Number: 3237 Maui BISH 1/15/1997
49 Dioscorea pentaphylla L. koolau range, kawailoa, kawaiiki ditch trail Moist wooded side gulch, under Aleurites and Metrosideros. PRESERVED_SPECIMEN st.john, h. collector number: 24704 Oahu BISH 10/14/1951
50 Dioscorea pentaphylla L. East Maui, Hana District, Hana Forest Reserve, Hana Ranch lands, mauka of Keakaamanu Alien forest slopes dominated by Psidium guajava and Ardisia elliptica. PRESERVED_SPECIMEN Imada, C.T. 2002-80 Maui BISH 11/11/2002
51 Dioscorea pentaphylla L. Halepuaa Forest Reserve in the Puna District PRESERVED_SPECIMEN Clarke, G. Collector Number: 79.205 Hawaii BISH 3/7/1979
52 Dioscorea pentaphylla L. wailau valley Climbing over shrubs. PRESERVED_SPECIMEN rock, j.f.c. collector number: 70221 Molokai BISH 4/20/1910
53 Dioscorea pentaphylla L. gulch west of ualapue Rocky woods. PRESERVED_SPECIMEN degener, o. collector number: 5562 Molokai BISH 6/30/1928
54 Dioscorea pentaphylla L. Koolau Range, Kawailoa, Kawaiiki Ditch trail Moist wooded side gulch, under Aleurites and Metrosideros. PRESERVED_SPECIMEN St. John, H. 24704 Oahu BISH 10/14/1951
55 Dioscorea pentaphylla L. East Maui, Hana District, 2.5 km W of Hana Hwy & Waikoloa Rd in the Kawaipapa area Tropical lowland forest planted w. introduced trees. PRESERVED_SPECIMEN Annable, C.R. 3237 Maui BISH 1/15/1997
56 Dioscorea pentaphylla L. East Maui, Hana District, Hana Forest Reserve, Hana Ranch lands, mauka of Keakaamanu Alien forest slopes dominated by Psidium guajava and Ardisia elliptica. PRESERVED_SPECIMEN Imada, C.T. LeGrande, M.; Jansen, S.; Crummer, J. Collector Number: 2002-80 Maui BISH 11/11/2002
57 Dioscorea pentaphylla L. Koolau Range; Kawailoa, Kawaiiki Ditch Trail, side gulch Rain forest. PRESERVED_SPECIMEN Wilson, K.A. 12 Oahu BISH 10/14/1951
58 Dioscorea pentaphylla L. koolau mts.; koolauloa, hauula, "hidden valley", above swanzy park.tch trail, side gulch Vine gowing on Pisonia PRESERVED_SPECIMEN teraoka, w. nagata, k.; whiteaker, l.; silva, b. collector number: 360 Oahu BISH 10/9/1980
59 Dioscorea pentaphylla L. Wainiha Valley, near dam PRESERVED_SPECIMEN Stone, B.C. 1505 Kauai BISH 12/26/1956
60 Dioscorea pentaphylla L. Kawailoa, Kawaiiki Ditch Trail PRESERVED_SPECIMEN St. John, H. 24834 Oahu BISH 7/13/1952
61 Dioscorea pentaphylla L. Manoa Valley, Waiakeakua, PRESERVED_SPECIMEN Wentworth, C.H. s.n. Oahu BISH 4/18/1905
62 Dioscorea pentaphylla L. ahupuaa of Halepuaa Forest Reserve in the Puna District. Found growing on the side of the old railroad On the side of the old railroad. PRESERVED_SPECIMEN ESP Field Crew Collector Number: s.n. Hawaii BISH 8/15/1979
63 Dioscorea pentaphylla L. Haupu, Kipu Moist forest. Climbing over trees. PRESERVED_SPECIMEN St. John, H. 13672 Kauai BISH 12/25/1933
64 Dioscorea pentaphylla L. palolo valley, kaau crater, frequent along trail on way to crater Along ditch in rice field PRESERVED_SPECIMEN egler, f.e. collector number: 37-309 Oahu BISH 7/29/1937
65 Dioscorea pentaphylla L. palolo, waiomao valley PRESERVED_SPECIMEN tongg, r. collector number: s.n. Molokai BISH 11/29/1942
66 Dioscorea pentaphylla L. Lihue-Koloa Forest Reserve; 3.2 miles NW of Wailua Experiment Station Vine climbing on Aleurites & Hibiscus overstory. In shade w/Zingiber, Nephrolepis, Oplismenus. PRESERVED_SPECIMEN Ishikawa, S. 277 Kauai BISH 8/20/1973
67 Dioscorea pentaphylla L. koolau range; kawailoa, kawaiiki ditch trail, side gulch Rain forest. PRESERVED_SPECIMEN wilson, k.a. collector number: 12 Oahu BISH 10/14/1951
68 Dioscorea pentaphylla L. Koolau Mts.; Koolauloa, Hauula, "Hidden Valley", above Swanzy Park.tch Trail, side gulch Vine gowing on Pisonia PRESERVED_SPECIMEN Teraoka, W. 360 Oahu BISH 10/9/1980
69 Dioscorea pentaphylla L. Palolo Valley, Kaau Crater, frequent along trail on way to crater Koa zone. PRESERVED_SPECIMEN Egler, F.E. 37-309 Oahu BISH 7/29/1937
70 Dioscorea pentaphylla L. Punaluu; Castle Trail Loose rocky soil, steep slope, ohia woods. PRESERVED_SPECIMEN St. John, H. 23376 Oahu BISH 5/23/1948
71 Dioscorea pentaphylla L. west niu gulch, at branch in stream Koa zone in opening in forest. PRESERVED_SPECIMEN egler, f.e. collector number: 37-195 Oahu BISH 7/20/1937
72 Dioscorea pentaphylla L. North Molokai; slopes of Waiahookalo Stream valley Steep, rocky slopes w/secondary forest of Psidium guajava, Syzygium malaccense & native remnants incl. Diospyros sandwicensis, Metrosideros & Nestegis w/understory of Kalanchoe pinnata & Zingiber zerumbet. Common along stream. PRESERVED_SPECIMEN Lorence, D.H. 5419 Molokai BISH 7/31/1987
73 Dioscorea pentaphylla L. manoa valley, waiakeakua, tubens PRESERVED_SPECIMEN wentworth, c.h. Oahu BISH 8/8/1935
74 Dioscorea pentaphylla L. Kapakahi Gulch Koa zone in ope kukui forest. PRESERVED_SPECIMEN Egler, F.E. 37-215 Oahu BISH 7/22/1937
75 Dioscorea pentaphylla L. Palolo Valley PRESERVED_SPECIMEN Forbes, C.N. 1968.O Oahu BISH 10/22/1914
76 Dioscorea pentaphylla L. West Niu Gulch, at branch in stream Koa zone in opening in forest. PRESERVED_SPECIMEN Egler, F.E. 37-195 Oahu BISH 7/20/1937
77 Dioscorea pentaphylla L. east maui, hana district, kalahu point, ptbg kahanu garden In remnant Pandanus forest SSE of Pi`ilanihale Heiau in rough a`a lava. Common throughout forest. PRESERVED_SPECIMEN flynn, t. collector number: 2487 Maui BISH 10/20/1987
78 Dioscorea pentaphylla L. wailau valley floor PRESERVED_SPECIMEN higashino, p.k. allen, m.s. collector number: 10374 Molokai BISH 8/13/1984
79 Dioscorea pentaphylla L. Halepuaa Forest Reserve in the Puna District PRESERVED_SPECIMEN Clarke, G. 79.205 Hawaii BISH 3/7/1979
80 Dioscorea pentaphylla L. Ewa Dist.; Waimalu Metrosideros forest. PRESERVED_SPECIMEN Grant, M.L. 7338 Oahu BISH 9/9/1934
81 Dioscorea pentaphylla L. wainiha valley, near dam wet; Metrosideros, Psychotria PRESERVED_SPECIMEN stone, b.c. collector number: 1505 Molokai BISH 12/26/1956
82 Dioscorea pentaphylla L. halepuaa forest reserve in the puna district PRESERVED_SPECIMEN clarke, g. collector number: 79.205 Hawaii BISH 3/7/1979
83 Dioscorea pentaphylla L. haupu, kipu Moist forest. Climbing over trees. PRESERVED_SPECIMEN st.john, h. fosberg, f.r. collector number: 13672 Kauai BISH 12/25/1933
84 Dioscorea pentaphylla L. 2.5km w of hana hwy & waikaloa rd, in the kawaipapa area Tropical lowland forest, planted with introduced trees PRESERVED_SPECIMEN c. r. annable 3237 NY 1/15/1997
85 Dioscorea pentaphylla L. 2.5km W of Hana Hwy & Waikaloa Rd, in the Kawaipapa area Tropical lowland forest, planted with introduced trees PRESERVED_SPECIMEN C. R. Annable 3237 NY 1/15/1997
86 Dioscorea pentaphylla L. Kalahu Point. PTBG Kahanu Garden. SSE of Pi'ilanihale Heiau on rough aa lava. Elev. 0-50 ft. In remnant Pandanus forest. PRESERVED_SPECIMEN Tim Flynn 2487 Maui PTBG 10/20/1987
87 Dioscorea pentaphylla L. Kealia Forest Reserve; Makaleha Ditch intake on Makaleha Stream. PRESERVED_SPECIMEN Tim Flynn 1332 Kauai PTBG 10/25/1985
88 Dioscorea pentaphylla L. West Maui, Honolua Valley. Lowland Wet Riparian Forest. PRESERVED_SPECIMEN Hank Oppenheimer H100604 Maui PTBG 10/4/2006
89 Dioscorea pentaphylla L. Kuilau Trail, below Keahua Arboretum. Disturbed lowland windward Forest PRESERVED_SPECIMEN Natalia Tangalin 3559 Kauai PTBG 11/22/2013
90 Dioscorea pentaphylla L. Slopes of Waiohookalo Stream Valley (N. Molokai); steep, rocky slopes. Alt. 100-180 m. With secondary forest. PRESERVED_SPECIMEN David H. Lorence 5419 Molokai PTBG 7/31/1987
91 Dioscorea pentaphylla L. waimalu, koolay range, oahu, territory of hawaii PRESERVED_SPECIMEN o. degener 10110 Hawaii US 11/3/1935
92 Dioscorea pentaphylla L. coast nw of hilo PRESERVED_SPECIMEN wilkes explor. exped. s.n. Hawaii US
93 Dioscorea pentaphylla L. Waimalu, Koolau Range, at middle elevation PRESERVED_SPECIMEN Degener, O.; Park; Potter; Shigeura; Takamoto, -. 10110 Oahu US 11/3/1935
94 Dioscorea pentaphylla L. honaunau PRESERVED_SPECIMEN a. s. hitchcock 14545 Hawaii US 9/3/1916
95 Dioscorea pentaphylla L. waimalu, koolau range, at middle elevation PRESERVED_SPECIMEN o. degener, park, potter, shigeura & -. takamoto 10110 Oahu US 11/3/1935
96 Dioscorea pentaphylla L. Honaunau PRESERVED_SPECIMEN Hitchcock, A. S. 14545 Hawaii US 9/3/2016
97 Dioscorea pentaphylla L. 3.2 mi NW of Wailua Experiment Station, Lihue-Koloa Forest Reserve PRESERVED_SPECIMEN Ishikawa, S. S. 277 Kauai US 8/20/1973
98 Dioscorea pentaphylla L. oahu PRESERVED_SPECIMEN st.john, h. collector number: 24834 Oahu BISH
99 Dioscorea pentaphylla L. PRESERVED_SPECIMEN degener, otto 10110 USCH 11/3/1935
100 Dioscorea pentaphylla L. Wailau Valley Climbing over shrubs. PRESERVED_SPECIMEN Rock, J.F.C. 70221 Molokai BISH 4/20/1910
101 Dioscorea pentaphylla L. waioli valley PRESERVED_SPECIMEN forbes, c.n. collector number: 91.k Kauai BISH 7/29/1909
102 Dioscorea pentaphylla L. palolo valley PRESERVED_SPECIMEN forbes, c.n. collector number: 1968.o Oahu BISH 10/22/1914
103 Dioscorea pentaphylla L. East Maui, Hana District, Kalahu Point, PTBG Kahanu Garden In remnant Pandanus forest SSE of Pi`ilanihale Heiau in rough a`a lava. Common throughout forest. PRESERVED_SPECIMEN Flynn, T. 2487 Maui BISH 10/20/1987
104 Dioscorea pentaphylla L. koolau range; waimalu Flowering only bare landslide at middle elevation. PRESERVED_SPECIMEN degener, o. park, k.k.; potter; shigeura; takamoto collector number: 10110 Oahu BISH 11/3/1935
105 Dioscorea pentaphylla L. 3.2 mi nw of wailua experiment station, lihue-koloa forest reserve PRESERVED_SPECIMEN s. s. ishikawa 277 Kauai US 8/20/1973
106 Dioscorea pentaphylla L. head of hanakoa valley Wet woods. PRESERVED_SPECIMEN st.john, h. collector number: 24968 Kauai BISH 1/3/1953
107 Dioscorea pentaphylla L. Koolau Range; Palolo, Waiomao Valley Intermediate moist. PRESERVED_SPECIMEN Nishida, T. s.n. Oahu BISH 10/5/1943
108 Dioscorea pentaphylla L. punaluu; castle trail Loose rocky soil, steep slope, ohia woods. PRESERVED_SPECIMEN st.john, h. collector number: 23376 Oahu BISH 5/23/1948
109 Dioscorea pentaphylla L. Wailau Valley floor PRESERVED_SPECIMEN Higashino, P.K. 10374 Molokai BISH 8/13/1984
110 Dioscorea pentaphylla L. wainiha; wainiha valley, bottom of small side gulch Moist lower forest. Climbing over trees. PRESERVED_SPECIMEN st.john, h. collector number: 13908 Kauai BISH 1/1/1934
111 Dioscorea pentaphylla L. ewa dist.; waimalu Metrosideros forest. PRESERVED_SPECIMEN grant, m.l. collector number: 7338 Oahu BISH 9/9/1934
112 Dioscorea pentaphylla L. Waimano Trail PRESERVED_SPECIMEN Kerr, M. 439-1 Oahu BISH 10/7/1945
113 Dioscorea pentaphylla L. Koolau Range; Palolo, Waiomao Valley Climbing over low shrubs, dense undergrowth. PRESERVED_SPECIMEN Kuykendall, J.R. 45 Oahu BISH 11/29/1942
114 Dioscorea pentaphylla L. Gulch west of Ualapue Rocky woods. PRESERVED_SPECIMEN Degener, O. 5562 Molokai BISH 6/30/1928
115 Dioscorea pentaphylla L. east maui, hana district, 2.5 km w of hana hwy & waikoloa rd in the kawaipapa area Tropical lowland forest planted w. introduced trees. PRESERVED_SPECIMEN annable, c.r. hobdy, r. collector number: 3237 Maui BISH 1/15/1997
116 Dioscorea pentaphylla L. waimano trail PRESERVED_SPECIMEN kerr, m. collector number: 439-1 Oahu BISH 10/7/1945
117 Dioscorea pentaphylla L. waimalu Damp shaded rocks in moderately dry mesic ohia forest w/ Aleurites and Eugenia malaccensis common in wetter gullies PRESERVED_SPECIMEN brown, f.b.h. collector number: 11282 Oahu BISH 12/31/1925
118 Dioscorea pentaphylla L. ahupuaa of halepuaa forest reserve in the puna district. found growing on the side of the old railroad On the side of the old railroad. PRESERVED_SPECIMEN esp field crew collector number: s.n. Hawaii BISH 8/15/1979
119 Dioscorea pentaphylla L. ahupuaa of Halepuaa Forest Reserve in the Puna District. Found growing on the side of the old railroad On the side of the old railroad. PRESERVED_SPECIMEN ESP Field Crew s.n. Hawaii BISH 8/15/1979
120 Dioscorea pentaphylla L. head of wailau valley Rain forest near banana patch. PRESERVED_SPECIMEN st.john, h. et al. collector number: 23531 Molokai BISH 12/27/1948
121 Dioscorea pentaphylla L. Wainiha; Wainiha Valley, bottom of small side gulch Moist lower forest. Climbing over trees. PRESERVED_SPECIMEN St. John, H. 13908 Kauai BISH 1/1/1934
122 Dioscorea pentaphylla L. Palolo, Waiomao Valley Open shrubby area in valley. PRESERVED_SPECIMEN Tongg, R. s.n. Oahu BISH 11/29/1942
123 Dioscorea pentaphylla L. north molokai; slopes of waiahookalo stream valley Steep, rocky slopes w/secondary forest of Psidium guajava, Syzygium malaccense & native remnants incl. Diospyros sandwicensis, Metrosideros & Nestegis w/understory of Kalanchoe pinnata & Zingiber zerumbet. Common along stream. PRESERVED_SPECIMEN lorence, d.h. wagner, w.l.; perlman, s.; montgomery, s. collector number: 5419 Molokai BISH 7/31/1987
124 Dioscorea pentaphylla L. Head of Wailau Valley Rain forest near banana patch. PRESERVED_SPECIMEN St. John, H. 23531 Molokai BISH 12/27/1948
125 Dioscorea pentaphylla L. koolau range; palolo, waiomao valley Intermediate moist. PRESERVED_SPECIMEN nishida, t. collector number: s.n. Oahu BISH 10/5/1943
126 Dioscorea pentaphylla L. kapakahi gulch Koa zone in ope kukui forest. PRESERVED_SPECIMEN egler, f.e. collector number: 37-215 Oahu BISH 7/22/1937
127 Dioscorea pentaphylla L. lihue-koloa forest reserve; 3.2 miles nw of wailua experiment station Vine climbing on Aleurites & Hibiscus overstory. In shade w/Zingiber, Nephrolepis, Oplismenus. PRESERVED_SPECIMEN ishikawa, s. collector number: 277 Kauai BISH 8/20/1973
128 Dioscorea pentaphylla L. east maui, hana district, hana forest reserve, hana ranch lands, mauka of keakaamanu Alien forest slopes dominated by Psidium guajava and Ardisia elliptica. PRESERVED_SPECIMEN imada, c.t. legrande, m.; jansen, s.; crummer, j. collector number: 2002-80 Maui BISH 11/11/2002
129 Dioscorea pentaphylla L. Waimalu PRESERVED_SPECIMEN Brown, F.B.H. 11282 Oahu BISH 12/31/1925
130 Dioscorea pentaphylla L. Waioli Valley PRESERVED_SPECIMEN Forbes, C.N. 91.K Kauai BISH 7/29/1909
131 Dioscorea pentaphylla L. ahupuaa of Halepuaa Forest Reserve in the Puna District. Found growing on the side of the old railroad Tropical lowland forest, planted with introduced trees PRESERVED_SPECIMEN C. R. Annable 3237 Hawaii HAPI 1/15/1997
132 Dioscorea pentaphylla L. koolau range; palolo, waiomao valley Climbing over low shrubs, dense undergrowth. PRESERVED_SPECIMEN kuykendall, j.r. collector number: 45 Oahu BISH 11/29/1942
133 Dioscorea pentaphylla L. Koolau Range; Waimalu Flowering only bare landslide at middle elevation. PRESERVED_SPECIMEN Degener, O. 10110 Oahu BISH 11/3/1935
134 Dioscorea pentaphylla L. East Maui, Hana District, Hana Forest Reserve, Hana Ranch lands, mauka of Keakaamanu Tropical lowland forest, planted with introduced trees UNKNOWN C. R. Annable Maui HAPI 1/15/1997
135 Dioscorea pentaphylla L. Head of Hanakoa Valley Wet woods. PRESERVED_SPECIMEN St. John, H. 24968 Kauai BISH 1/3/1953
136 Dioscorea pentaphylla L. East Maui, Hana District, 2.5 km W of Hana Hwy & Waikoloa Rd in the Kawaipapa area Tropical lowland forest planted w. introduced trees. PRESERVED_SPECIMEN Annable, C.R. Hobdy, R. Collector Number: 3237 Maui BISH 1/15/1997
137 Dioscorea pentaphylla L. koolau range, kawailoa, kawaiiki ditch trail Moist wooded side gulch, under Aleurites and Metrosideros. PRESERVED_SPECIMEN st.john, h. collector number: 24704 Oahu BISH 10/14/1951
138 Dioscorea pentaphylla L. East Maui, Hana District, Hana Forest Reserve, Hana Ranch lands, mauka of Keakaamanu Alien forest slopes dominated by Psidium guajava and Ardisia elliptica. PRESERVED_SPECIMEN Imada, C.T. 2002-80 Maui BISH 11/11/2002
139 Dioscorea pentaphylla L. Halepuaa Forest Reserve in the Puna District PRESERVED_SPECIMEN Clarke, G. Collector Number: 79.205 Hawaii BISH 3/7/1979
140 Dioscorea pentaphylla L. wailau valley Climbing over shrubs. PRESERVED_SPECIMEN rock, j.f.c. collector number: 70221 Molokai BISH 4/20/1910
141 Dioscorea pentaphylla L. gulch west of ualapue Rocky woods. PRESERVED_SPECIMEN degener, o. collector number: 5562 Molokai BISH 6/30/1928
142 Dioscorea pentaphylla L. Koolau Range, Kawailoa, Kawaiiki Ditch trail Moist wooded side gulch, under Aleurites and Metrosideros. PRESERVED_SPECIMEN St. John, H. 24704 Oahu BISH 10/14/1951
143 Dioscorea pentaphylla L. East Maui, Hana District, 2.5 km W of Hana Hwy & Waikoloa Rd in the Kawaipapa area Tropical lowland forest planted w. introduced trees. PRESERVED_SPECIMEN Annable, C.R. 3237 Maui BISH 1/15/1997
144 Dioscorea pentaphylla L. East Maui, Hana District, Hana Forest Reserve, Hana Ranch lands, mauka of Keakaamanu Alien forest slopes dominated by Psidium guajava and Ardisia elliptica. PRESERVED_SPECIMEN Imada, C.T. LeGrande, M.; Jansen, S.; Crummer, J. Collector Number: 2002-80 Maui BISH 11/11/2002
145 Dioscorea pentaphylla L. Koolau Range; Kawailoa, Kawaiiki Ditch Trail, side gulch Rain forest. PRESERVED_SPECIMEN Wilson, K.A. 12 Oahu BISH 10/14/1951
146 Dioscorea pentaphylla L. koolau mts.; koolauloa, hauula, "hidden valley", above swanzy park.tch trail, side gulch Vine gowing on Pisonia PRESERVED_SPECIMEN teraoka, w. nagata, k.; whiteaker, l.; silva, b. collector number: 360 Oahu BISH 10/9/1980
147 Dioscorea pentaphylla L. Wainiha Valley, near dam PRESERVED_SPECIMEN Stone, B.C. 1505 Kauai BISH 12/26/1956
148 Dioscorea pentaphylla L. Kawailoa, Kawaiiki Ditch Trail PRESERVED_SPECIMEN St. John, H. 24834 Oahu BISH 7/13/1952
149 Dioscorea pentaphylla L. Manoa Valley, Waiakeakua, PRESERVED_SPECIMEN Wentworth, C.H. s.n. Oahu BISH 4/18/1905
150 Dioscorea pentaphylla L. ahupuaa of Halepuaa Forest Reserve in the Puna District. Found growing on the side of the old railroad On the side of the old railroad. PRESERVED_SPECIMEN ESP Field Crew Collector Number: s.n. Hawaii BISH 8/15/1979
151 Dioscorea pentaphylla L. Haupu, Kipu Moist forest. Climbing over trees. PRESERVED_SPECIMEN St. John, H. 13672 Kauai BISH 12/25/1933
152 Dioscorea pentaphylla L. palolo valley, kaau crater, frequent along trail on way to crater Along ditch in rice field PRESERVED_SPECIMEN egler, f.e. collector number: 37-309 Oahu BISH 7/29/1937
153 Dioscorea pentaphylla L. palolo, waiomao valley PRESERVED_SPECIMEN tongg, r. collector number: s.n. Molokai BISH 11/29/1942
154 Dioscorea pentaphylla L. Lihue-Koloa Forest Reserve; 3.2 miles NW of Wailua Experiment Station Vine climbing on Aleurites & Hibiscus overstory. In shade w/Zingiber, Nephrolepis, Oplismenus. PRESERVED_SPECIMEN Ishikawa, S. 277 Kauai BISH 8/20/1973
155 Dioscorea pentaphylla L. koolau range; kawailoa, kawaiiki ditch trail, side gulch Rain forest. PRESERVED_SPECIMEN wilson, k.a. collector number: 12 Oahu BISH 10/14/1951
156 Dioscorea pentaphylla L. Koolau Mts.; Koolauloa, Hauula, "Hidden Valley", above Swanzy Park.tch Trail, side gulch Vine gowing on Pisonia PRESERVED_SPECIMEN Teraoka, W. 360 Oahu BISH 10/9/1980
157 Dioscorea pentaphylla L. Palolo Valley, Kaau Crater, frequent along trail on way to crater Koa zone. PRESERVED_SPECIMEN Egler, F.E. 37-309 Oahu BISH 7/29/1937
158 Dioscorea pentaphylla L. Punaluu; Castle Trail Loose rocky soil, steep slope, ohia woods. PRESERVED_SPECIMEN St. John, H. 23376 Oahu BISH 5/23/1948
159 Dioscorea pentaphylla L. west niu gulch, at branch in stream Koa zone in opening in forest. PRESERVED_SPECIMEN egler, f.e. collector number: 37-195 Oahu BISH 7/20/1937
160 Dioscorea pentaphylla L. North Molokai; slopes of Waiahookalo Stream valley Steep, rocky slopes w/secondary forest of Psidium guajava, Syzygium malaccense & native remnants incl. Diospyros sandwicensis, Metrosideros & Nestegis w/understory of Kalanchoe pinnata & Zingiber zerumbet. Common along stream. PRESERVED_SPECIMEN Lorence, D.H. 5419 Molokai BISH 7/31/1987
161 Dioscorea pentaphylla L. manoa valley, waiakeakua, tubens PRESERVED_SPECIMEN wentworth, c.h. Oahu BISH 8/8/1935
162 Dioscorea pentaphylla L. Kapakahi Gulch Koa zone in ope kukui forest. PRESERVED_SPECIMEN Egler, F.E. 37-215 Oahu BISH 7/22/1937
163 Dioscorea pentaphylla L. Palolo Valley PRESERVED_SPECIMEN Forbes, C.N. 1968.O Oahu BISH 10/22/1914
164 Dioscorea pentaphylla L. West Niu Gulch, at branch in stream Koa zone in opening in forest. PRESERVED_SPECIMEN Egler, F.E. 37-195 Oahu BISH 7/20/1937
165 Dioscorea pentaphylla L. east maui, hana district, kalahu point, ptbg kahanu garden In remnant Pandanus forest SSE of Pi`ilanihale Heiau in rough a`a lava. Common throughout forest. PRESERVED_SPECIMEN flynn, t. collector number: 2487 Maui BISH 10/20/1987
166 Dioscorea pentaphylla L. wailau valley floor PRESERVED_SPECIMEN higashino, p.k. allen, m.s. collector number: 10374 Molokai BISH 8/13/1984
167 Dioscorea pentaphylla L. Halepuaa Forest Reserve in the Puna District PRESERVED_SPECIMEN Clarke, G. 79.205 Hawaii BISH 3/7/1979
168 Dioscorea pentaphylla L. Ewa Dist.; Waimalu Metrosideros forest. PRESERVED_SPECIMEN Grant, M.L. 7338 Oahu BISH 9/9/1934
169 Dioscorea pentaphylla L. wainiha valley, near dam wet; Metrosideros, Psychotria PRESERVED_SPECIMEN stone, b.c. collector number: 1505 Molokai BISH 12/26/1956
170 Dioscorea pentaphylla L. halepuaa forest reserve in the puna district PRESERVED_SPECIMEN clarke, g. collector number: 79.205 Hawaii BISH 3/7/1979
171 Dioscorea pentaphylla L. haupu, kipu Moist forest. Climbing over trees. PRESERVED_SPECIMEN st.john, h. fosberg, f.r. collector number: 13672 Kauai BISH 12/25/1933
172 Dioscorea pentaphylla L. 2.5km w of hana hwy & waikaloa rd, in the kawaipapa area Tropical lowland forest, planted with introduced trees PRESERVED_SPECIMEN c. r. annable 3237 NY 1/15/1997
173 Dioscorea pentaphylla L. 2.5km W of Hana Hwy & Waikaloa Rd, in the Kawaipapa area Tropical lowland forest, planted with introduced trees PRESERVED_SPECIMEN C. R. Annable 3237 NY 1/15/1997
174 Dioscorea pentaphylla L. Kalahu Point. PTBG Kahanu Garden. SSE of Pi'ilanihale Heiau on rough aa lava. Elev. 0-50 ft. In remnant Pandanus forest. PRESERVED_SPECIMEN Tim Flynn 2487 Maui PTBG 10/20/1987
175 Dioscorea pentaphylla L. Kealia Forest Reserve; Makaleha Ditch intake on Makaleha Stream. PRESERVED_SPECIMEN Tim Flynn 1332 Kauai PTBG 10/25/1985
176 Dioscorea pentaphylla L. West Maui, Honolua Valley. Lowland Wet Riparian Forest. PRESERVED_SPECIMEN Hank Oppenheimer H100604 Maui PTBG 10/4/2006
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