Dioscorea pentaphylla

L. (1753)

This name is accepted

Kingdom: Viridiplantae Phylum: Magnoliophyta Class/Clade: Monocots Order: Dioscoreales Family: Dioscoreaceae Genus: Dioscorea

fiveleaf yam, pi‘a, pi‘a [pia] Hawai‘i

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Description

Key Characters:

Growth Form: Vines.

Stems: Stems twining to the left, = angled, up to 10 m long, pilose to hirsute, glabrate with age, usually with numerous scattered prickles, tubers nearly always solitary, variable in shape, growing near the surface or over 1 m underground.

Roots: Fibrous root system.

Leaves: Leaves compound (palmate) or rarely simple. Alternate. Leaflets 3–5, elliptic or oblong, 5–10 cm long, 2–4 cm wide, the central one oblanceolate or obovate, the lateral ones somewhat asymmetrical. Apex obtuse or acute to acuminate. Surfaces with yellowish brown pubescent at least on lower surface, axils with bulbils and occasionally bearing globose to horseshoe shaped tubers ca. 1 cm in diameter. Leaflet margins entire. Petioles 5-11 cm long, densely reddish brown pubescent. Stipules absent.

Flowers: Staminate flowers in axillary, spike-like racemes up to 30 cm long, near the ends of the branches, reddish pubescent, pistillate flowers in long pendent spikes 5–25 cm long. Flowers unisexual, the plants dioecious. Calyx of staminate flowers with Tepals broadly lanceolate, ca. 1 mm long. Calyx of pistillate flowers with tepals ca. 1.5 mm long. Staminate flower with 3 stamen and 3 staminodes (longer than stamens); pistillate flowers with 6 rudimentary staminodes. Ovary inferior, villous, 3-carpellate.

Fruit: Capsules oblong; 20–25 mm long; glabrous or glabrate; shallowly emarginate at both ends; the wings up to 6 mm wide. Seeds winged on basal side only.

Ploidy: 2n = 40; 70; 80; 144

Habitat: Naturalized primarily in windward mesic valleys.

Elevation Range: 115–325 m.

Historical Distribution

Images

Uses and Culture

USES

  • Tubers eaten in times of famine (Krauss 1993:11,12).

Natural History

Statewide Status

Naturalized

Island Status

Kaua'i Naturalized
O'ahu Naturalized
Molokai Naturalized
Maui Naturalized
Hawai'i Naturalized

Dispersal Agents


Pollinators

Bibliography

Name Published In: Sp. Pl.: 1032 (1753)

Other References

Wagner et al. 1990:1437 (K, O, Mo, M, H)

Occurrences

SNo. Scientific Name Scientific Name Authorship Locality Habitat Basis of Record Recorded By Record Number Island Source Date
1 Dioscorea pentaphylla L. waimalu, koolau range, at middle elevation PRESERVED_SPECIMEN o. degener, park, potter, shigeura & -. takamoto 10110 Oahu US 11/3/1935
2 Dioscorea pentaphylla L. 3.2 mi NW of Wailua Experiment Station, Lihue-Koloa Forest Reserve PRESERVED_SPECIMEN Ishikawa, S. S. 277 Kauai US 8/20/1973
3 Dioscorea pentaphylla L. Honaunau PRESERVED_SPECIMEN Hitchcock, A. S. 14545 Hawaii US 9/3/2016
4 Dioscorea pentaphylla L. oahu PRESERVED_SPECIMEN st.john, h. collector number: 24834 Oahu BISH
5 Dioscorea pentaphylla L. PRESERVED_SPECIMEN degener, otto 10110 USCH 11/3/1935
6 Dioscorea pentaphylla L. Wailau Valley Climbing over shrubs. PRESERVED_SPECIMEN Rock, J.F.C. 70221 Molokai BISH 4/20/1910
7 Dioscorea pentaphylla L. waioli valley PRESERVED_SPECIMEN forbes, c.n. collector number: 91.k Kauai BISH 7/29/1909
8 Dioscorea pentaphylla L. palolo valley PRESERVED_SPECIMEN forbes, c.n. collector number: 1968.o Oahu BISH 10/22/1914
9 Dioscorea pentaphylla L. East Maui, Hana District, Kalahu Point, PTBG Kahanu Garden In remnant Pandanus forest SSE of Pi`ilanihale Heiau in rough a`a lava. Common throughout forest. PRESERVED_SPECIMEN Flynn, T. 2487 Maui BISH 10/20/1987
10 Dioscorea pentaphylla L. koolau range; waimalu Flowering only bare landslide at middle elevation. PRESERVED_SPECIMEN degener, o. park, k.k.; potter; shigeura; takamoto collector number: 10110 Oahu BISH 11/3/1935
11 Dioscorea pentaphylla L. 3.2 mi nw of wailua experiment station, lihue-koloa forest reserve PRESERVED_SPECIMEN s. s. ishikawa 277 Kauai US 8/20/1973
12 Dioscorea pentaphylla L. head of hanakoa valley Wet woods. PRESERVED_SPECIMEN st.john, h. collector number: 24968 Kauai BISH 1/3/1953
13 Dioscorea pentaphylla L. Koolau Range; Palolo, Waiomao Valley Intermediate moist. PRESERVED_SPECIMEN Nishida, T. s.n. Oahu BISH 10/5/1943
14 Dioscorea pentaphylla L. Wailau Valley floor PRESERVED_SPECIMEN Higashino, P.K. 10374 Molokai BISH 8/13/1984
15 Dioscorea pentaphylla L. punaluu; castle trail Loose rocky soil, steep slope, ohia woods. PRESERVED_SPECIMEN st.john, h. collector number: 23376 Oahu BISH 5/23/1948
16 Dioscorea pentaphylla L. Waimano Trail PRESERVED_SPECIMEN Kerr, M. 439-1 Oahu BISH 10/7/1945
17 Dioscorea pentaphylla L. ewa dist.; waimalu Metrosideros forest. PRESERVED_SPECIMEN grant, m.l. collector number: 7338 Oahu BISH 9/9/1934
18 Dioscorea pentaphylla L. Koolau Range; Palolo, Waiomao Valley Climbing over low shrubs, dense undergrowth. PRESERVED_SPECIMEN Kuykendall, J.R. 45 Oahu BISH 11/29/1942
19 Dioscorea pentaphylla L. Gulch west of Ualapue Rocky woods. PRESERVED_SPECIMEN Degener, O. 5562 Molokai BISH 6/30/1928
20 Dioscorea pentaphylla L. ahupuaa of halepuaa forest reserve in the puna district. found growing on the side of the old railroad On the side of the old railroad. PRESERVED_SPECIMEN esp field crew collector number: s.n. Hawaii BISH 8/15/1979
21 Dioscorea pentaphylla L. wainiha; wainiha valley, bottom of small side gulch Moist lower forest. Climbing over trees. PRESERVED_SPECIMEN st.john, h. collector number: 13908 Kauai BISH 1/1/1934
22 Dioscorea pentaphylla L. waimano trail PRESERVED_SPECIMEN kerr, m. collector number: 439-1 Oahu BISH 10/7/1945
23 Dioscorea pentaphylla L. ahupuaa of Halepuaa Forest Reserve in the Puna District. Found growing on the side of the old railroad On the side of the old railroad. PRESERVED_SPECIMEN ESP Field Crew s.n. Hawaii BISH 8/15/1979
24 Dioscorea pentaphylla L. east maui, hana district, 2.5 km w of hana hwy & waikoloa rd in the kawaipapa area Tropical lowland forest planted w. introduced trees. PRESERVED_SPECIMEN annable, c.r. hobdy, r. collector number: 3237 Maui BISH 1/15/1997
25 Dioscorea pentaphylla L. waimalu Damp shaded rocks in moderately dry mesic ohia forest w/ Aleurites and Eugenia malaccensis common in wetter gullies PRESERVED_SPECIMEN brown, f.b.h. collector number: 11282 Oahu BISH 12/31/1925
26 Dioscorea pentaphylla L. head of wailau valley Rain forest near banana patch. PRESERVED_SPECIMEN st.john, h. et al. collector number: 23531 Molokai BISH 12/27/1948
27 Dioscorea pentaphylla L. Palolo, Waiomao Valley Open shrubby area in valley. PRESERVED_SPECIMEN Tongg, R. s.n. Oahu BISH 11/29/1942
28 Dioscorea pentaphylla L. north molokai; slopes of waiahookalo stream valley Steep, rocky slopes w/secondary forest of Psidium guajava, Syzygium malaccense & native remnants incl. Diospyros sandwicensis, Metrosideros & Nestegis w/understory of Kalanchoe pinnata & Zingiber zerumbet. Common along stream. PRESERVED_SPECIMEN lorence, d.h. wagner, w.l.; perlman, s.; montgomery, s. collector number: 5419 Molokai BISH 7/31/1987
29 Dioscorea pentaphylla L. Wainiha; Wainiha Valley, bottom of small side gulch Moist lower forest. Climbing over trees. PRESERVED_SPECIMEN St. John, H. 13908 Kauai BISH 1/1/1934
30 Dioscorea pentaphylla L. koolau range; palolo, waiomao valley Intermediate moist. PRESERVED_SPECIMEN nishida, t. collector number: s.n. Oahu BISH 10/5/1943
31 Dioscorea pentaphylla L. Head of Wailau Valley Rain forest near banana patch. PRESERVED_SPECIMEN St. John, H. 23531 Molokai BISH 12/27/1948
32 Dioscorea pentaphylla L. kapakahi gulch Koa zone in ope kukui forest. PRESERVED_SPECIMEN egler, f.e. collector number: 37-215 Oahu BISH 7/22/1937
33 Dioscorea pentaphylla L. gulch west of ualapue Rocky woods. PRESERVED_SPECIMEN degener, o. collector number: 5562 Molokai BISH 6/30/1928
34 Dioscorea pentaphylla L. koolau range; palolo, waiomao valley Climbing over low shrubs, dense undergrowth. PRESERVED_SPECIMEN kuykendall, j.r. collector number: 45 Oahu BISH 11/29/1942
35 Dioscorea pentaphylla L. lihue-koloa forest reserve; 3.2 miles nw of wailua experiment station Vine climbing on Aleurites & Hibiscus overstory. In shade w/Zingiber, Nephrolepis, Oplismenus. PRESERVED_SPECIMEN ishikawa, s. collector number: 277 Kauai BISH 8/20/1973
36 Dioscorea pentaphylla L. east maui, hana district, hana forest reserve, hana ranch lands, mauka of keakaamanu Alien forest slopes dominated by Psidium guajava and Ardisia elliptica. PRESERVED_SPECIMEN imada, c.t. legrande, m.; jansen, s.; crummer, j. collector number: 2002-80 Maui BISH 11/11/2002
37 Dioscorea pentaphylla L. Waioli Valley PRESERVED_SPECIMEN Forbes, C.N. 91.K Kauai BISH 7/29/1909
38 Dioscorea pentaphylla L. Waimalu PRESERVED_SPECIMEN Brown, F.B.H. 11282 Oahu BISH 12/31/1925
39 Dioscorea pentaphylla L. ahupuaa of Halepuaa Forest Reserve in the Puna District. Found growing on the side of the old railroad Tropical lowland forest, planted with introduced trees PRESERVED_SPECIMEN C. R. Annable 3237 Hawaii HAPI 1/15/1997
40 Dioscorea pentaphylla L. East Maui, Hana District, Hana Forest Reserve, Hana Ranch lands, mauka of Keakaamanu Tropical lowland forest, planted with introduced trees UNKNOWN C. R. Annable Maui HAPI 1/15/1997
41 Dioscorea pentaphylla L. Koolau Range; Waimalu Flowering only bare landslide at middle elevation. PRESERVED_SPECIMEN Degener, O. 10110 Oahu BISH 11/3/1935
42 Dioscorea pentaphylla L. Head of Hanakoa Valley Wet woods. PRESERVED_SPECIMEN St. John, H. 24968 Kauai BISH 1/3/1953
43 Dioscorea pentaphylla L. koolau range, kawailoa, kawaiiki ditch trail Moist wooded side gulch, under Aleurites and Metrosideros. PRESERVED_SPECIMEN st.john, h. collector number: 24704 Oahu BISH 10/14/1951
44 Dioscorea pentaphylla L. East Maui, Hana District, 2.5 km W of Hana Hwy & Waikoloa Rd in the Kawaipapa area Tropical lowland forest planted w. introduced trees. PRESERVED_SPECIMEN Annable, C.R. Hobdy, R. Collector Number: 3237 Maui BISH 1/15/1997
45 Dioscorea pentaphylla L. East Maui, Hana District, Hana Forest Reserve, Hana Ranch lands, mauka of Keakaamanu Alien forest slopes dominated by Psidium guajava and Ardisia elliptica. PRESERVED_SPECIMEN Imada, C.T. 2002-80 Maui BISH 11/11/2002
46 Dioscorea pentaphylla L. Halepuaa Forest Reserve in the Puna District PRESERVED_SPECIMEN Clarke, G. Collector Number: 79.205 Hawaii BISH 3/7/1979
47 Dioscorea pentaphylla L. wailau valley Climbing over shrubs. PRESERVED_SPECIMEN rock, j.f.c. collector number: 70221 Molokai BISH 4/20/1910
48 Dioscorea pentaphylla L. Koolau Range, Kawailoa, Kawaiiki Ditch trail Moist wooded side gulch, under Aleurites and Metrosideros. PRESERVED_SPECIMEN St. John, H. 24704 Oahu BISH 10/14/1951
49 Dioscorea pentaphylla L. East Maui, Hana District, 2.5 km W of Hana Hwy & Waikoloa Rd in the Kawaipapa area Tropical lowland forest planted w. introduced trees. PRESERVED_SPECIMEN Annable, C.R. 3237 Maui BISH 1/15/1997
50 Dioscorea pentaphylla L. East Maui, Hana District, Hana Forest Reserve, Hana Ranch lands, mauka of Keakaamanu Alien forest slopes dominated by Psidium guajava and Ardisia elliptica. PRESERVED_SPECIMEN Imada, C.T. LeGrande, M.; Jansen, S.; Crummer, J. Collector Number: 2002-80 Maui BISH 11/11/2002
51 Dioscorea pentaphylla L. Koolau Range; Kawailoa, Kawaiiki Ditch Trail, side gulch Rain forest. PRESERVED_SPECIMEN Wilson, K.A. 12 Oahu BISH 10/14/1951
52 Dioscorea pentaphylla L. koolau mts.; koolauloa, hauula, "hidden valley", above swanzy park.tch trail, side gulch Vine gowing on Pisonia PRESERVED_SPECIMEN teraoka, w. nagata, k.; whiteaker, l.; silva, b. collector number: 360 Oahu BISH 10/9/1980
53 Dioscorea pentaphylla L. Wainiha Valley, near dam PRESERVED_SPECIMEN Stone, B.C. 1505 Kauai BISH 12/26/1956
54 Dioscorea pentaphylla L. Kawailoa, Kawaiiki Ditch Trail PRESERVED_SPECIMEN St. John, H. 24834 Oahu BISH 7/13/1952
55 Dioscorea pentaphylla L. ahupuaa of Halepuaa Forest Reserve in the Puna District. Found growing on the side of the old railroad On the side of the old railroad. PRESERVED_SPECIMEN ESP Field Crew Collector Number: s.n. Hawaii BISH 8/15/1979
56 Dioscorea pentaphylla L. Manoa Valley, Waiakeakua, PRESERVED_SPECIMEN Wentworth, C.H. s.n. Oahu BISH 4/18/1905
57 Dioscorea pentaphylla L. Haupu, Kipu Moist forest. Climbing over trees. PRESERVED_SPECIMEN St. John, H. 13672 Kauai BISH 12/25/1933
58 Dioscorea pentaphylla L. palolo, waiomao valley PRESERVED_SPECIMEN tongg, r. collector number: s.n. Molokai BISH 11/29/1942
59 Dioscorea pentaphylla L. palolo valley, kaau crater, frequent along trail on way to crater Along ditch in rice field PRESERVED_SPECIMEN egler, f.e. collector number: 37-309 Oahu BISH 7/29/1937
60 Dioscorea pentaphylla L. koolau range; kawailoa, kawaiiki ditch trail, side gulch Rain forest. PRESERVED_SPECIMEN wilson, k.a. collector number: 12 Oahu BISH 10/14/1951
61 Dioscorea pentaphylla L. Koolau Mts.; Koolauloa, Hauula, "Hidden Valley", above Swanzy Park.tch Trail, side gulch Vine gowing on Pisonia PRESERVED_SPECIMEN Teraoka, W. 360 Oahu BISH 10/9/1980
62 Dioscorea pentaphylla L. Lihue-Koloa Forest Reserve; 3.2 miles NW of Wailua Experiment Station Vine climbing on Aleurites & Hibiscus overstory. In shade w/Zingiber, Nephrolepis, Oplismenus. PRESERVED_SPECIMEN Ishikawa, S. 277 Kauai BISH 8/20/1973
63 Dioscorea pentaphylla L. Punaluu; Castle Trail Loose rocky soil, steep slope, ohia woods. PRESERVED_SPECIMEN St. John, H. 23376 Oahu BISH 5/23/1948
64 Dioscorea pentaphylla L. west niu gulch, at branch in stream Koa zone in opening in forest. PRESERVED_SPECIMEN egler, f.e. collector number: 37-195 Oahu BISH 7/20/1937
65 Dioscorea pentaphylla L. North Molokai; slopes of Waiahookalo Stream valley Steep, rocky slopes w/secondary forest of Psidium guajava, Syzygium malaccense & native remnants incl. Diospyros sandwicensis, Metrosideros & Nestegis w/understory of Kalanchoe pinnata & Zingiber zerumbet. Common along stream. PRESERVED_SPECIMEN Lorence, D.H. 5419 Molokai BISH 7/31/1987
66 Dioscorea pentaphylla L. 2.5km w of hana hwy & waikaloa rd, in the kawaipapa area Tropical lowland forest, planted with introduced trees PRESERVED_SPECIMEN c. r. annable 3237 NY 1/15/1997
67 Dioscorea pentaphylla L. manoa valley, waiakeakua, tubens PRESERVED_SPECIMEN wentworth, c.h. Oahu BISH 8/8/1935
68 Dioscorea pentaphylla L. Palolo Valley, Kaau Crater, frequent along trail on way to crater Koa zone. PRESERVED_SPECIMEN Egler, F.E. 37-309 Oahu BISH 7/29/1937
69 Dioscorea pentaphylla L. Kapakahi Gulch Koa zone in ope kukui forest. PRESERVED_SPECIMEN Egler, F.E. 37-215 Oahu BISH 7/22/1937
70 Dioscorea pentaphylla L. Palolo Valley PRESERVED_SPECIMEN Forbes, C.N. 1968.O Oahu BISH 10/22/1914
71 Dioscorea pentaphylla L. West Niu Gulch, at branch in stream Koa zone in opening in forest. PRESERVED_SPECIMEN Egler, F.E. 37-195 Oahu BISH 7/20/1937
72 Dioscorea pentaphylla L. wailau valley floor PRESERVED_SPECIMEN higashino, p.k. allen, m.s. collector number: 10374 Molokai BISH 8/13/1984
73 Dioscorea pentaphylla L. east maui, hana district, kalahu point, ptbg kahanu garden In remnant Pandanus forest SSE of Pi`ilanihale Heiau in rough a`a lava. Common throughout forest. PRESERVED_SPECIMEN flynn, t. collector number: 2487 Maui BISH 10/20/1987
74 Dioscorea pentaphylla L. Halepuaa Forest Reserve in the Puna District PRESERVED_SPECIMEN Clarke, G. 79.205 Hawaii BISH 3/7/1979
75 Dioscorea pentaphylla L. wainiha valley, near dam wet; Metrosideros, Psychotria PRESERVED_SPECIMEN stone, b.c. collector number: 1505 Molokai BISH 12/26/1956
76 Dioscorea pentaphylla L. Ewa Dist.; Waimalu Metrosideros forest. PRESERVED_SPECIMEN Grant, M.L. 7338 Oahu BISH 9/9/1934
77 Dioscorea pentaphylla L. halepuaa forest reserve in the puna district PRESERVED_SPECIMEN clarke, g. collector number: 79.205 Hawaii BISH 3/7/1979
78 Dioscorea pentaphylla L. haupu, kipu Moist forest. Climbing over trees. PRESERVED_SPECIMEN st.john, h. fosberg, f.r. collector number: 13672 Kauai BISH 12/25/1933
79 Dioscorea pentaphylla L. 2.5km W of Hana Hwy & Waikaloa Rd, in the Kawaipapa area Tropical lowland forest, planted with introduced trees PRESERVED_SPECIMEN C. R. Annable 3237 NY 1/15/1997
80 Dioscorea pentaphylla L. Kalahu Point. PTBG Kahanu Garden. SSE of Pi'ilanihale Heiau on rough aa lava. Elev. 0-50 ft. In remnant Pandanus forest. PRESERVED_SPECIMEN Tim Flynn 2487 Maui PTBG 10/20/1987
81 Dioscorea pentaphylla L. Kealia Forest Reserve; Makaleha Ditch intake on Makaleha Stream. PRESERVED_SPECIMEN Tim Flynn 1332 Kauai PTBG 10/25/1985
82 Dioscorea pentaphylla L. coast nw of hilo PRESERVED_SPECIMEN wilkes explor. exped. s.n. Hawaii US
83 Dioscorea pentaphylla L. West Maui, Honolua Valley. Lowland Wet Riparian Forest. PRESERVED_SPECIMEN Hank Oppenheimer H100604 Maui PTBG 10/4/2006
84 Dioscorea pentaphylla L. Kuilau Trail, below Keahua Arboretum. Disturbed lowland windward Forest PRESERVED_SPECIMEN Natalia Tangalin 3559 Kauai PTBG 11/22/2013
85 Dioscorea pentaphylla L. Slopes of Waiohookalo Stream Valley (N. Molokai); steep, rocky slopes. Alt. 100-180 m. With secondary forest. PRESERVED_SPECIMEN David H. Lorence 5419 Molokai PTBG 7/31/1987
86 Dioscorea pentaphylla L. waimalu, koolay range, oahu, territory of hawaii PRESERVED_SPECIMEN o. degener 10110 Hawaii US 11/3/1935
87 Dioscorea pentaphylla L. honaunau PRESERVED_SPECIMEN a. s. hitchcock 14545 Hawaii US 9/3/1916
88 Dioscorea pentaphylla L. Waimalu, Koolau Range, at middle elevation PRESERVED_SPECIMEN Degener, O.; Park; Potter; Shigeura; Takamoto, -. 10110 Oahu US 11/3/1935
89 Dioscorea pentaphylla L. waimalu, koolau range, at middle elevation PRESERVED_SPECIMEN o. degener, park, potter, shigeura & -. takamoto 10110 Oahu US 11/3/1935
90 Dioscorea pentaphylla L. 3.2 mi NW of Wailua Experiment Station, Lihue-Koloa Forest Reserve PRESERVED_SPECIMEN Ishikawa, S. S. 277 Kauai US 8/20/1973
91 Dioscorea pentaphylla L. Honaunau PRESERVED_SPECIMEN Hitchcock, A. S. 14545 Hawaii US 9/3/2016
92 Dioscorea pentaphylla L. oahu PRESERVED_SPECIMEN st.john, h. collector number: 24834 Oahu BISH
93 Dioscorea pentaphylla L. PRESERVED_SPECIMEN degener, otto 10110 USCH 11/3/1935
94 Dioscorea pentaphylla L. Wailau Valley Climbing over shrubs. PRESERVED_SPECIMEN Rock, J.F.C. 70221 Molokai BISH 4/20/1910
95 Dioscorea pentaphylla L. waioli valley PRESERVED_SPECIMEN forbes, c.n. collector number: 91.k Kauai BISH 7/29/1909
96 Dioscorea pentaphylla L. palolo valley PRESERVED_SPECIMEN forbes, c.n. collector number: 1968.o Oahu BISH 10/22/1914
97 Dioscorea pentaphylla L. East Maui, Hana District, Kalahu Point, PTBG Kahanu Garden In remnant Pandanus forest SSE of Pi`ilanihale Heiau in rough a`a lava. Common throughout forest. PRESERVED_SPECIMEN Flynn, T. 2487 Maui BISH 10/20/1987
98 Dioscorea pentaphylla L. koolau range; waimalu Flowering only bare landslide at middle elevation. PRESERVED_SPECIMEN degener, o. park, k.k.; potter; shigeura; takamoto collector number: 10110 Oahu BISH 11/3/1935
99 Dioscorea pentaphylla L. 3.2 mi nw of wailua experiment station, lihue-koloa forest reserve PRESERVED_SPECIMEN s. s. ishikawa 277 Kauai US 8/20/1973
100 Dioscorea pentaphylla L. head of hanakoa valley Wet woods. PRESERVED_SPECIMEN st.john, h. collector number: 24968 Kauai BISH 1/3/1953
101 Dioscorea pentaphylla L. Koolau Range; Palolo, Waiomao Valley Intermediate moist. PRESERVED_SPECIMEN Nishida, T. s.n. Oahu BISH 10/5/1943
102 Dioscorea pentaphylla L. Wailau Valley floor PRESERVED_SPECIMEN Higashino, P.K. 10374 Molokai BISH 8/13/1984
103 Dioscorea pentaphylla L. punaluu; castle trail Loose rocky soil, steep slope, ohia woods. PRESERVED_SPECIMEN st.john, h. collector number: 23376 Oahu BISH 5/23/1948
104 Dioscorea pentaphylla L. Waimano Trail PRESERVED_SPECIMEN Kerr, M. 439-1 Oahu BISH 10/7/1945
105 Dioscorea pentaphylla L. ewa dist.; waimalu Metrosideros forest. PRESERVED_SPECIMEN grant, m.l. collector number: 7338 Oahu BISH 9/9/1934
106 Dioscorea pentaphylla L. Koolau Range; Palolo, Waiomao Valley Climbing over low shrubs, dense undergrowth. PRESERVED_SPECIMEN Kuykendall, J.R. 45 Oahu BISH 11/29/1942
107 Dioscorea pentaphylla L. Gulch west of Ualapue Rocky woods. PRESERVED_SPECIMEN Degener, O. 5562 Molokai BISH 6/30/1928
108 Dioscorea pentaphylla L. ahupuaa of halepuaa forest reserve in the puna district. found growing on the side of the old railroad On the side of the old railroad. PRESERVED_SPECIMEN esp field crew collector number: s.n. Hawaii BISH 8/15/1979
109 Dioscorea pentaphylla L. wainiha; wainiha valley, bottom of small side gulch Moist lower forest. Climbing over trees. PRESERVED_SPECIMEN st.john, h. collector number: 13908 Kauai BISH 1/1/1934
110 Dioscorea pentaphylla L. waimano trail PRESERVED_SPECIMEN kerr, m. collector number: 439-1 Oahu BISH 10/7/1945
111 Dioscorea pentaphylla L. ahupuaa of Halepuaa Forest Reserve in the Puna District. Found growing on the side of the old railroad On the side of the old railroad. PRESERVED_SPECIMEN ESP Field Crew s.n. Hawaii BISH 8/15/1979
112 Dioscorea pentaphylla L. east maui, hana district, 2.5 km w of hana hwy & waikoloa rd in the kawaipapa area Tropical lowland forest planted w. introduced trees. PRESERVED_SPECIMEN annable, c.r. hobdy, r. collector number: 3237 Maui BISH 1/15/1997
113 Dioscorea pentaphylla L. waimalu Damp shaded rocks in moderately dry mesic ohia forest w/ Aleurites and Eugenia malaccensis common in wetter gullies PRESERVED_SPECIMEN brown, f.b.h. collector number: 11282 Oahu BISH 12/31/1925
114 Dioscorea pentaphylla L. head of wailau valley Rain forest near banana patch. PRESERVED_SPECIMEN st.john, h. et al. collector number: 23531 Molokai BISH 12/27/1948
115 Dioscorea pentaphylla L. Palolo, Waiomao Valley Open shrubby area in valley. PRESERVED_SPECIMEN Tongg, R. s.n. Oahu BISH 11/29/1942
116 Dioscorea pentaphylla L. north molokai; slopes of waiahookalo stream valley Steep, rocky slopes w/secondary forest of Psidium guajava, Syzygium malaccense & native remnants incl. Diospyros sandwicensis, Metrosideros & Nestegis w/understory of Kalanchoe pinnata & Zingiber zerumbet. Common along stream. PRESERVED_SPECIMEN lorence, d.h. wagner, w.l.; perlman, s.; montgomery, s. collector number: 5419 Molokai BISH 7/31/1987
117 Dioscorea pentaphylla L. Wainiha; Wainiha Valley, bottom of small side gulch Moist lower forest. Climbing over trees. PRESERVED_SPECIMEN St. John, H. 13908 Kauai BISH 1/1/1934
118 Dioscorea pentaphylla L. koolau range; palolo, waiomao valley Intermediate moist. PRESERVED_SPECIMEN nishida, t. collector number: s.n. Oahu BISH 10/5/1943
119 Dioscorea pentaphylla L. Head of Wailau Valley Rain forest near banana patch. PRESERVED_SPECIMEN St. John, H. 23531 Molokai BISH 12/27/1948
120 Dioscorea pentaphylla L. kapakahi gulch Koa zone in ope kukui forest. PRESERVED_SPECIMEN egler, f.e. collector number: 37-215 Oahu BISH 7/22/1937
121 Dioscorea pentaphylla L. gulch west of ualapue Rocky woods. PRESERVED_SPECIMEN degener, o. collector number: 5562 Molokai BISH 6/30/1928
122 Dioscorea pentaphylla L. koolau range; palolo, waiomao valley Climbing over low shrubs, dense undergrowth. PRESERVED_SPECIMEN kuykendall, j.r. collector number: 45 Oahu BISH 11/29/1942
123 Dioscorea pentaphylla L. lihue-koloa forest reserve; 3.2 miles nw of wailua experiment station Vine climbing on Aleurites & Hibiscus overstory. In shade w/Zingiber, Nephrolepis, Oplismenus. PRESERVED_SPECIMEN ishikawa, s. collector number: 277 Kauai BISH 8/20/1973
124 Dioscorea pentaphylla L. east maui, hana district, hana forest reserve, hana ranch lands, mauka of keakaamanu Alien forest slopes dominated by Psidium guajava and Ardisia elliptica. PRESERVED_SPECIMEN imada, c.t. legrande, m.; jansen, s.; crummer, j. collector number: 2002-80 Maui BISH 11/11/2002
125 Dioscorea pentaphylla L. Waioli Valley PRESERVED_SPECIMEN Forbes, C.N. 91.K Kauai BISH 7/29/1909
126 Dioscorea pentaphylla L. Waimalu PRESERVED_SPECIMEN Brown, F.B.H. 11282 Oahu BISH 12/31/1925
127 Dioscorea pentaphylla L. ahupuaa of Halepuaa Forest Reserve in the Puna District. Found growing on the side of the old railroad Tropical lowland forest, planted with introduced trees PRESERVED_SPECIMEN C. R. Annable 3237 Hawaii HAPI 1/15/1997
128 Dioscorea pentaphylla L. East Maui, Hana District, Hana Forest Reserve, Hana Ranch lands, mauka of Keakaamanu Tropical lowland forest, planted with introduced trees UNKNOWN C. R. Annable Maui HAPI 1/15/1997
129 Dioscorea pentaphylla L. Koolau Range; Waimalu Flowering only bare landslide at middle elevation. PRESERVED_SPECIMEN Degener, O. 10110 Oahu BISH 11/3/1935
130 Dioscorea pentaphylla L. Head of Hanakoa Valley Wet woods. PRESERVED_SPECIMEN St. John, H. 24968 Kauai BISH 1/3/1953
131 Dioscorea pentaphylla L. koolau range, kawailoa, kawaiiki ditch trail Moist wooded side gulch, under Aleurites and Metrosideros. PRESERVED_SPECIMEN st.john, h. collector number: 24704 Oahu BISH 10/14/1951
132 Dioscorea pentaphylla L. East Maui, Hana District, 2.5 km W of Hana Hwy & Waikoloa Rd in the Kawaipapa area Tropical lowland forest planted w. introduced trees. PRESERVED_SPECIMEN Annable, C.R. Hobdy, R. Collector Number: 3237 Maui BISH 1/15/1997
133 Dioscorea pentaphylla L. East Maui, Hana District, Hana Forest Reserve, Hana Ranch lands, mauka of Keakaamanu Alien forest slopes dominated by Psidium guajava and Ardisia elliptica. PRESERVED_SPECIMEN Imada, C.T. 2002-80 Maui BISH 11/11/2002
134 Dioscorea pentaphylla L. Halepuaa Forest Reserve in the Puna District PRESERVED_SPECIMEN Clarke, G. Collector Number: 79.205 Hawaii BISH 3/7/1979
135 Dioscorea pentaphylla L. wailau valley Climbing over shrubs. PRESERVED_SPECIMEN rock, j.f.c. collector number: 70221 Molokai BISH 4/20/1910
136 Dioscorea pentaphylla L. Koolau Range, Kawailoa, Kawaiiki Ditch trail Moist wooded side gulch, under Aleurites and Metrosideros. PRESERVED_SPECIMEN St. John, H. 24704 Oahu BISH 10/14/1951
137 Dioscorea pentaphylla L. East Maui, Hana District, 2.5 km W of Hana Hwy & Waikoloa Rd in the Kawaipapa area Tropical lowland forest planted w. introduced trees. PRESERVED_SPECIMEN Annable, C.R. 3237 Maui BISH 1/15/1997
138 Dioscorea pentaphylla L. East Maui, Hana District, Hana Forest Reserve, Hana Ranch lands, mauka of Keakaamanu Alien forest slopes dominated by Psidium guajava and Ardisia elliptica. PRESERVED_SPECIMEN Imada, C.T. LeGrande, M.; Jansen, S.; Crummer, J. Collector Number: 2002-80 Maui BISH 11/11/2002
139 Dioscorea pentaphylla L. Koolau Range; Kawailoa, Kawaiiki Ditch Trail, side gulch Rain forest. PRESERVED_SPECIMEN Wilson, K.A. 12 Oahu BISH 10/14/1951
140 Dioscorea pentaphylla L. koolau mts.; koolauloa, hauula, "hidden valley", above swanzy park.tch trail, side gulch Vine gowing on Pisonia PRESERVED_SPECIMEN teraoka, w. nagata, k.; whiteaker, l.; silva, b. collector number: 360 Oahu BISH 10/9/1980
141 Dioscorea pentaphylla L. Wainiha Valley, near dam PRESERVED_SPECIMEN Stone, B.C. 1505 Kauai BISH 12/26/1956
142 Dioscorea pentaphylla L. Kawailoa, Kawaiiki Ditch Trail PRESERVED_SPECIMEN St. John, H. 24834 Oahu BISH 7/13/1952
143 Dioscorea pentaphylla L. ahupuaa of Halepuaa Forest Reserve in the Puna District. Found growing on the side of the old railroad On the side of the old railroad. PRESERVED_SPECIMEN ESP Field Crew Collector Number: s.n. Hawaii BISH 8/15/1979
144 Dioscorea pentaphylla L. Manoa Valley, Waiakeakua, PRESERVED_SPECIMEN Wentworth, C.H. s.n. Oahu BISH 4/18/1905
145 Dioscorea pentaphylla L. Haupu, Kipu Moist forest. Climbing over trees. PRESERVED_SPECIMEN St. John, H. 13672 Kauai BISH 12/25/1933
146 Dioscorea pentaphylla L. palolo, waiomao valley PRESERVED_SPECIMEN tongg, r. collector number: s.n. Molokai BISH 11/29/1942
147 Dioscorea pentaphylla L. palolo valley, kaau crater, frequent along trail on way to crater Along ditch in rice field PRESERVED_SPECIMEN egler, f.e. collector number: 37-309 Oahu BISH 7/29/1937
148 Dioscorea pentaphylla L. koolau range; kawailoa, kawaiiki ditch trail, side gulch Rain forest. PRESERVED_SPECIMEN wilson, k.a. collector number: 12 Oahu BISH 10/14/1951
149 Dioscorea pentaphylla L. Koolau Mts.; Koolauloa, Hauula, "Hidden Valley", above Swanzy Park.tch Trail, side gulch Vine gowing on Pisonia PRESERVED_SPECIMEN Teraoka, W. 360 Oahu BISH 10/9/1980
150 Dioscorea pentaphylla L. Lihue-Koloa Forest Reserve; 3.2 miles NW of Wailua Experiment Station Vine climbing on Aleurites & Hibiscus overstory. In shade w/Zingiber, Nephrolepis, Oplismenus. PRESERVED_SPECIMEN Ishikawa, S. 277 Kauai BISH 8/20/1973
151 Dioscorea pentaphylla L. Punaluu; Castle Trail Loose rocky soil, steep slope, ohia woods. PRESERVED_SPECIMEN St. John, H. 23376 Oahu BISH 5/23/1948
152 Dioscorea pentaphylla L. west niu gulch, at branch in stream Koa zone in opening in forest. PRESERVED_SPECIMEN egler, f.e. collector number: 37-195 Oahu BISH 7/20/1937
153 Dioscorea pentaphylla L. North Molokai; slopes of Waiahookalo Stream valley Steep, rocky slopes w/secondary forest of Psidium guajava, Syzygium malaccense & native remnants incl. Diospyros sandwicensis, Metrosideros & Nestegis w/understory of Kalanchoe pinnata & Zingiber zerumbet. Common along stream. PRESERVED_SPECIMEN Lorence, D.H. 5419 Molokai BISH 7/31/1987
154 Dioscorea pentaphylla L. 2.5km w of hana hwy & waikaloa rd, in the kawaipapa area Tropical lowland forest, planted with introduced trees PRESERVED_SPECIMEN c. r. annable 3237 NY 1/15/1997
155 Dioscorea pentaphylla L. manoa valley, waiakeakua, tubens PRESERVED_SPECIMEN wentworth, c.h. Oahu BISH 8/8/1935
156 Dioscorea pentaphylla L. Palolo Valley, Kaau Crater, frequent along trail on way to crater Koa zone. PRESERVED_SPECIMEN Egler, F.E. 37-309 Oahu BISH 7/29/1937
157 Dioscorea pentaphylla L. Kapakahi Gulch Koa zone in ope kukui forest. PRESERVED_SPECIMEN Egler, F.E. 37-215 Oahu BISH 7/22/1937
158 Dioscorea pentaphylla L. Palolo Valley PRESERVED_SPECIMEN Forbes, C.N. 1968.O Oahu BISH 10/22/1914
159 Dioscorea pentaphylla L. West Niu Gulch, at branch in stream Koa zone in opening in forest. PRESERVED_SPECIMEN Egler, F.E. 37-195 Oahu BISH 7/20/1937
160 Dioscorea pentaphylla L. wailau valley floor PRESERVED_SPECIMEN higashino, p.k. allen, m.s. collector number: 10374 Molokai BISH 8/13/1984
161 Dioscorea pentaphylla L. east maui, hana district, kalahu point, ptbg kahanu garden In remnant Pandanus forest SSE of Pi`ilanihale Heiau in rough a`a lava. Common throughout forest. PRESERVED_SPECIMEN flynn, t. collector number: 2487 Maui BISH 10/20/1987
162 Dioscorea pentaphylla L. Halepuaa Forest Reserve in the Puna District PRESERVED_SPECIMEN Clarke, G. 79.205 Hawaii BISH 3/7/1979
163 Dioscorea pentaphylla L. wainiha valley, near dam wet; Metrosideros, Psychotria PRESERVED_SPECIMEN stone, b.c. collector number: 1505 Molokai BISH 12/26/1956
164 Dioscorea pentaphylla L. Ewa Dist.; Waimalu Metrosideros forest. PRESERVED_SPECIMEN Grant, M.L. 7338 Oahu BISH 9/9/1934
165 Dioscorea pentaphylla L. halepuaa forest reserve in the puna district PRESERVED_SPECIMEN clarke, g. collector number: 79.205 Hawaii BISH 3/7/1979
166 Dioscorea pentaphylla L. haupu, kipu Moist forest. Climbing over trees. PRESERVED_SPECIMEN st.john, h. fosberg, f.r. collector number: 13672 Kauai BISH 12/25/1933
167 Dioscorea pentaphylla L. 2.5km W of Hana Hwy & Waikaloa Rd, in the Kawaipapa area Tropical lowland forest, planted with introduced trees PRESERVED_SPECIMEN C. R. Annable 3237 NY 1/15/1997
168 Dioscorea pentaphylla L. Kalahu Point. PTBG Kahanu Garden. SSE of Pi'ilanihale Heiau on rough aa lava. Elev. 0-50 ft. In remnant Pandanus forest. PRESERVED_SPECIMEN Tim Flynn 2487 Maui PTBG 10/20/1987
169 Dioscorea pentaphylla L. Kealia Forest Reserve; Makaleha Ditch intake on Makaleha Stream. PRESERVED_SPECIMEN Tim Flynn 1332 Kauai PTBG 10/25/1985
170 Dioscorea pentaphylla L. coast nw of hilo PRESERVED_SPECIMEN wilkes explor. exped. s.n. Hawaii US
171 Dioscorea pentaphylla L. West Maui, Honolua Valley. Lowland Wet Riparian Forest. PRESERVED_SPECIMEN Hank Oppenheimer H100604 Maui PTBG 10/4/2006
172 Dioscorea pentaphylla L. Kuilau Trail, below Keahua Arboretum. Disturbed lowland windward Forest PRESERVED_SPECIMEN Natalia Tangalin 3559 Kauai PTBG 11/22/2013
173 Dioscorea pentaphylla L. Slopes of Waiohookalo Stream Valley (N. Molokai); steep, rocky slopes. Alt. 100-180 m. With secondary forest. PRESERVED_SPECIMEN David H. Lorence 5419 Molokai PTBG 7/31/1987
174 Dioscorea pentaphylla L. waimalu, koolay range, oahu, territory of hawaii PRESERVED_SPECIMEN o. degener 10110 Hawaii US 11/3/1935
175 Dioscorea pentaphylla L. honaunau PRESERVED_SPECIMEN a. s. hitchcock 14545 Hawaii US 9/3/1916
176 Dioscorea pentaphylla L. Waimalu, Koolau Range, at middle elevation PRESERVED_SPECIMEN Degener, O.; Park; Potter; Shigeura; Takamoto, -. 10110 Oahu US 11/3/1935