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Dioscorea pentaphylla

L. (1753)

This name is accepted

Kingdom: Viridiplantae Phylum: Magnoliophyta Class/Clade: Monocots Order: Dioscoreales Family: Dioscoreaceae Genus: Dioscorea

fiveleaf yam, pi‘a [pia] Hawai‘i

Description

Key Characters:

Growth Form: Vines.

Stems: Stems twining to the left, = angled, up to 10 m long, pilose to hirsute, glabrate with age, usually with numerous scattered prickles, tubers nearly always solitary, variable in shape, growing near the surface or over 1 m underground.

Roots: Fibrous root system.

Leaves: Leaves compound (palmate) or rarely simple. Alternate. Leaflets 3–5, elliptic or oblong, 5–10 cm long, 2–4 cm wide, the central one oblanceolate or obovate, the lateral ones somewhat asymmetrical. Apex obtuse or acute to acuminate. Surfaces with yellowish brown pubescent at least on lower surface, axils with bulbils and occasionally bearing globose to horseshoe shaped tubers ca. 1 cm in diameter. Leaflet margins entire. Petioles 5-11 cm long, densely reddish brown pubescent. Stipules absent.

Flowers: Staminate flowers in axillary, spike-like racemes up to 30 cm long, near the ends of the branches, reddish pubescent, pistillate flowers in long pendent spikes 5–25 cm long. Flowers unisexual, the plants dioecious. Calyx of staminate flowers with Tepals broadly lanceolate, ca. 1 mm long. Calyx of pistillate flowers with tepals ca. 1.5 mm long. Staminate flower with 3 stamen and 3 staminodes (longer than stamens); pistillate flowers with 6 rudimentary staminodes. Ovary inferior, villous, 3-carpellate.

Fruit: Capsules oblong; 20–25 mm long; glabrous or glabrate; shallowly emarginate at both ends; the wings up to 6 mm wide. Seeds winged on basal side only.

Ploidy: 2n = 40; 70; 80; 144

Habitat: Naturalized primarily in windward mesic valleys.

Elevation Range: 115–325 m.

Historical Distribution

Uses and Culture

USES

  • Tubers eaten in times of famine (Krauss 1993:11,12).

Natural History

Statewide Status

Naturalized

Island Status

Kaua'i Naturalized
O'ahu Naturalized
Molokai Naturalized
Maui Naturalized
Hawai'i Naturalized

Dispersal Agents


Pollinators

Specimens

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Bibliography

Name Published In: Sp. Pl.: 1032 (1753)

Other References

Wagner et al. 1990:1437 (K, O, Mo, M, H)

Occurrences

SNo. Scientific Name Scientific Name Authorship Locality Habitat Basis of Record Recorded By Record Number Island Source Date
1 Dioscorea pentaphylla L. Kalahu Point. PTBG Kahanu Garden. SSE of Pi'ilanihale Heiau on rough aa lava. Elev. 0-50 ft. In remnant Pandanus forest. PRESERVED_SPECIMEN Tim Flynn 2487 Maui PTBG 10/20/1987
2 Dioscorea pentaphylla L. Kealia Forest Reserve; Makaleha Ditch intake on Makaleha Stream. PRESERVED_SPECIMEN Tim Flynn 1332 Kauai PTBG 10/25/1985
3 Dioscorea pentaphylla L. West Maui, Honolua Valley. Lowland Wet Riparian Forest. PRESERVED_SPECIMEN Hank Oppenheimer H100604 Maui PTBG 10/4/2006
4 Dioscorea pentaphylla L. Kuilau Trail, below Keahua Arboretum. Disturbed lowland windward Forest PRESERVED_SPECIMEN Natalia Tangalin 3559 Kauai PTBG 11/22/2013
5 Dioscorea pentaphylla L. Waimalu, Koolau Range, at middle elevation PRESERVED_SPECIMEN Degener, O.; Park; Potter; Shigeura; Takamoto, -. 10110 Oahu US 11/3/1935
6 Dioscorea pentaphylla L. Slopes of Waiohookalo Stream Valley (N. Molokai); steep, rocky slopes. Alt. 100-180 m. With secondary forest. PRESERVED_SPECIMEN David H. Lorence 5419 Molokai PTBG 7/31/1987
7 Dioscorea pentaphylla L. waimalu, koolay range, oahu, territory of hawaii PRESERVED_SPECIMEN o. degener 10110 Hawaii US 11/3/1935
8 Dioscorea pentaphylla L. coast nw of hilo PRESERVED_SPECIMEN wilkes explor. exped. s.n. Hawaii US
9 Dioscorea pentaphylla L. honaunau PRESERVED_SPECIMEN a. s. hitchcock 14545 Hawaii US 9/3/1916
10 Dioscorea pentaphylla L. waimalu, koolau range, at middle elevation PRESERVED_SPECIMEN o. degener, park, potter, shigeura & -. takamoto 10110 Oahu US 11/3/1935
11 Dioscorea pentaphylla L. 3.2 mi NW of Wailua Experiment Station, Lihue-Koloa Forest Reserve PRESERVED_SPECIMEN Ishikawa, S. S. 277 Kauai US 8/20/1973
12 Dioscorea pentaphylla L. Honaunau PRESERVED_SPECIMEN Hitchcock, A. S. 14545 Hawaii US 9/3/2016
13 Dioscorea pentaphylla L. oahu PRESERVED_SPECIMEN st.john, h. collector number: 24834 Oahu BISH
14 Dioscorea pentaphylla L. PRESERVED_SPECIMEN degener, otto 10110 USCH 11/3/1935
15 Dioscorea pentaphylla L. Wailau Valley Climbing over shrubs. PRESERVED_SPECIMEN Rock, J.F.C. 70221 Molokai BISH 4/20/1910
16 Dioscorea pentaphylla L. waioli valley PRESERVED_SPECIMEN forbes, c.n. collector number: 91.k Kauai BISH 7/29/1909
17 Dioscorea pentaphylla L. East Maui, Hana District, Kalahu Point, PTBG Kahanu Garden In remnant Pandanus forest SSE of Pi`ilanihale Heiau in rough a`a lava. Common throughout forest. PRESERVED_SPECIMEN Flynn, T. 2487 Maui BISH 10/20/1987
18 Dioscorea pentaphylla L. koolau range; waimalu Flowering only bare landslide at middle elevation. PRESERVED_SPECIMEN degener, o. park, k.k.; potter; shigeura; takamoto collector number: 10110 Oahu BISH 11/3/1935
19 Dioscorea pentaphylla L. palolo valley PRESERVED_SPECIMEN forbes, c.n. collector number: 1968.o Oahu BISH 10/22/1914
20 Dioscorea pentaphylla L. wainiha; wainiha valley, bottom of small side gulch Moist lower forest. Climbing over trees. PRESERVED_SPECIMEN st.john, h. collector number: 13908 Kauai BISH 1/1/1934
21 Dioscorea pentaphylla L. head of hanakoa valley Wet woods. PRESERVED_SPECIMEN st.john, h. collector number: 24968 Kauai BISH 1/3/1953
22 Dioscorea pentaphylla L. 3.2 mi nw of wailua experiment station, lihue-koloa forest reserve PRESERVED_SPECIMEN s. s. ishikawa 277 Kauai US 8/20/1973
23 Dioscorea pentaphylla L. Wailau Valley floor PRESERVED_SPECIMEN Higashino, P.K. 10374 Molokai BISH 8/13/1984
24 Dioscorea pentaphylla L. Waimano Trail PRESERVED_SPECIMEN Kerr, M. 439-1 Oahu BISH 10/7/1945
25 Dioscorea pentaphylla L. Koolau Range; Palolo, Waiomao Valley Climbing over low shrubs, dense undergrowth. PRESERVED_SPECIMEN Kuykendall, J.R. 45 Oahu BISH 11/29/1942
26 Dioscorea pentaphylla L. Gulch west of Ualapue Rocky woods. PRESERVED_SPECIMEN Degener, O. 5562 Molokai BISH 6/30/1928
27 Dioscorea pentaphylla L. Koolau Range; Palolo, Waiomao Valley Intermediate moist. PRESERVED_SPECIMEN Nishida, T. s.n. Oahu BISH 10/5/1943
28 Dioscorea pentaphylla L. punaluu; castle trail Loose rocky soil, steep slope, ohia woods. PRESERVED_SPECIMEN st.john, h. collector number: 23376 Oahu BISH 5/23/1948
29 Dioscorea pentaphylla L. ewa dist.; waimalu Metrosideros forest. PRESERVED_SPECIMEN grant, m.l. collector number: 7338 Oahu BISH 9/9/1934
30 Dioscorea pentaphylla L. east maui, hana district, 2.5 km w of hana hwy & waikoloa rd in the kawaipapa area Tropical lowland forest planted w. introduced trees. PRESERVED_SPECIMEN annable, c.r. hobdy, r. collector number: 3237 Maui BISH 1/15/1997
31 Dioscorea pentaphylla L. waimalu Damp shaded rocks in moderately dry mesic ohia forest w/ Aleurites and Eugenia malaccensis common in wetter gullies PRESERVED_SPECIMEN brown, f.b.h. collector number: 11282 Oahu BISH 12/31/1925
32 Dioscorea pentaphylla L. lihue-koloa forest reserve; 3.2 miles nw of wailua experiment station Vine climbing on Aleurites & Hibiscus overstory. In shade w/Zingiber, Nephrolepis, Oplismenus. PRESERVED_SPECIMEN ishikawa, s. collector number: 277 Kauai BISH 8/20/1973
33 Dioscorea pentaphylla L. waimano trail PRESERVED_SPECIMEN kerr, m. collector number: 439-1 Oahu BISH 10/7/1945
34 Dioscorea pentaphylla L. ahupuaa of Halepuaa Forest Reserve in the Puna District. Found growing on the side of the old railroad On the side of the old railroad. PRESERVED_SPECIMEN ESP Field Crew s.n. Hawaii BISH 8/15/1979
35 Dioscorea pentaphylla L. head of wailau valley Rain forest near banana patch. PRESERVED_SPECIMEN st.john, h. et al. collector number: 23531 Molokai BISH 12/27/1948
36 Dioscorea pentaphylla L. Palolo, Waiomao Valley Open shrubby area in valley. PRESERVED_SPECIMEN Tongg, R. s.n. Oahu BISH 11/29/1942
37 Dioscorea pentaphylla L. north molokai; slopes of waiahookalo stream valley Steep, rocky slopes w/secondary forest of Psidium guajava, Syzygium malaccense & native remnants incl. Diospyros sandwicensis, Metrosideros & Nestegis w/understory of Kalanchoe pinnata & Zingiber zerumbet. Common along stream. PRESERVED_SPECIMEN lorence, d.h. wagner, w.l.; perlman, s.; montgomery, s. collector number: 5419 Molokai BISH 7/31/1987
38 Dioscorea pentaphylla L. koolau range; palolo, waiomao valley Intermediate moist. PRESERVED_SPECIMEN nishida, t. collector number: s.n. Oahu BISH 10/5/1943
39 Dioscorea pentaphylla L. ahupuaa of halepuaa forest reserve in the puna district. found growing on the side of the old railroad On the side of the old railroad. PRESERVED_SPECIMEN esp field crew collector number: s.n. Hawaii BISH 8/15/1979
40 Dioscorea pentaphylla L. Wainiha; Wainiha Valley, bottom of small side gulch Moist lower forest. Climbing over trees. PRESERVED_SPECIMEN St. John, H. 13908 Kauai BISH 1/1/1934
41 Dioscorea pentaphylla L. Head of Wailau Valley Rain forest near banana patch. PRESERVED_SPECIMEN St. John, H. 23531 Molokai BISH 12/27/1948
42 Dioscorea pentaphylla L. kapakahi gulch Koa zone in ope kukui forest. PRESERVED_SPECIMEN egler, f.e. collector number: 37-215 Oahu BISH 7/22/1937
43 Dioscorea pentaphylla L. east maui, hana district, hana forest reserve, hana ranch lands, mauka of keakaamanu Alien forest slopes dominated by Psidium guajava and Ardisia elliptica. PRESERVED_SPECIMEN imada, c.t. legrande, m.; jansen, s.; crummer, j. collector number: 2002-80 Maui BISH 11/11/2002
44 Dioscorea pentaphylla L. Waioli Valley PRESERVED_SPECIMEN Forbes, C.N. 91.K Kauai BISH 7/29/1909
45 Dioscorea pentaphylla L. Waimalu PRESERVED_SPECIMEN Brown, F.B.H. 11282 Oahu BISH 12/31/1925
46 Dioscorea pentaphylla L. koolau range; palolo, waiomao valley Climbing over low shrubs, dense undergrowth. PRESERVED_SPECIMEN kuykendall, j.r. collector number: 45 Oahu BISH 11/29/1942
47 Dioscorea pentaphylla L. Koolau Range; Kawailoa, Kawaiiki Ditch Trail, side gulch Rain forest. PRESERVED_SPECIMEN Wilson, K.A. 12 Oahu BISH 10/14/1951
48 Dioscorea pentaphylla L. ahupuaa of Halepuaa Forest Reserve in the Puna District. Found growing on the side of the old railroad Tropical lowland forest, planted with introduced trees PRESERVED_SPECIMEN C. R. Annable 3237 Hawaii HAPI 1/15/1997
49 Dioscorea pentaphylla L. wailau valley Climbing over shrubs. PRESERVED_SPECIMEN rock, j.f.c. collector number: 70221 Molokai BISH 4/20/1910
50 Dioscorea pentaphylla L. gulch west of ualapue Rocky woods. PRESERVED_SPECIMEN degener, o. collector number: 5562 Molokai BISH 6/30/1928
51 Dioscorea pentaphylla L. East Maui, Hana District, 2.5 km W of Hana Hwy & Waikoloa Rd in the Kawaipapa area Tropical lowland forest planted w. introduced trees. PRESERVED_SPECIMEN Annable, C.R. 3237 Maui BISH 1/15/1997
52 Dioscorea pentaphylla L. East Maui, Hana District, Hana Forest Reserve, Hana Ranch lands, mauka of Keakaamanu Tropical lowland forest, planted with introduced trees UNKNOWN C. R. Annable Maui HAPI 1/15/1997
53 Dioscorea pentaphylla L. Koolau Range; Waimalu Flowering only bare landslide at middle elevation. PRESERVED_SPECIMEN Degener, O. 10110 Oahu BISH 11/3/1935
54 Dioscorea pentaphylla L. Head of Hanakoa Valley Wet woods. PRESERVED_SPECIMEN St. John, H. 24968 Kauai BISH 1/3/1953
55 Dioscorea pentaphylla L. koolau range, kawailoa, kawaiiki ditch trail Moist wooded side gulch, under Aleurites and Metrosideros. PRESERVED_SPECIMEN st.john, h. collector number: 24704 Oahu BISH 10/14/1951
56 Dioscorea pentaphylla L. East Maui, Hana District, 2.5 km W of Hana Hwy & Waikoloa Rd in the Kawaipapa area Tropical lowland forest planted w. introduced trees. PRESERVED_SPECIMEN Annable, C.R. Hobdy, R. Collector Number: 3237 Maui BISH 1/15/1997
57 Dioscorea pentaphylla L. East Maui, Hana District, Hana Forest Reserve, Hana Ranch lands, mauka of Keakaamanu Alien forest slopes dominated by Psidium guajava and Ardisia elliptica. PRESERVED_SPECIMEN Imada, C.T. 2002-80 Maui BISH 11/11/2002
58 Dioscorea pentaphylla L. Halepuaa Forest Reserve in the Puna District PRESERVED_SPECIMEN Clarke, G. Collector Number: 79.205 Hawaii BISH 3/7/1979
59 Dioscorea pentaphylla L. Koolau Range, Kawailoa, Kawaiiki Ditch trail Moist wooded side gulch, under Aleurites and Metrosideros. PRESERVED_SPECIMEN St. John, H. 24704 Oahu BISH 10/14/1951
60 Dioscorea pentaphylla L. East Maui, Hana District, Hana Forest Reserve, Hana Ranch lands, mauka of Keakaamanu Alien forest slopes dominated by Psidium guajava and Ardisia elliptica. PRESERVED_SPECIMEN Imada, C.T. LeGrande, M.; Jansen, S.; Crummer, J. Collector Number: 2002-80 Maui BISH 11/11/2002
61 Dioscorea pentaphylla L. Wainiha Valley, near dam PRESERVED_SPECIMEN Stone, B.C. 1505 Kauai BISH 12/26/1956
62 Dioscorea pentaphylla L. Kawailoa, Kawaiiki Ditch Trail PRESERVED_SPECIMEN St. John, H. 24834 Oahu BISH 7/13/1952
63 Dioscorea pentaphylla L. koolau mts.; koolauloa, hauula, "hidden valley", above swanzy park.tch trail, side gulch Vine gowing on Pisonia PRESERVED_SPECIMEN teraoka, w. nagata, k.; whiteaker, l.; silva, b. collector number: 360 Oahu BISH 10/9/1980
64 Dioscorea pentaphylla L. ahupuaa of Halepuaa Forest Reserve in the Puna District. Found growing on the side of the old railroad On the side of the old railroad. PRESERVED_SPECIMEN ESP Field Crew Collector Number: s.n. Hawaii BISH 8/15/1979
65 Dioscorea pentaphylla L. Manoa Valley, Waiakeakua, PRESERVED_SPECIMEN Wentworth, C.H. s.n. Oahu BISH 4/18/1905
66 Dioscorea pentaphylla L. Haupu, Kipu Moist forest. Climbing over trees. PRESERVED_SPECIMEN St. John, H. 13672 Kauai BISH 12/25/1933
67 Dioscorea pentaphylla L. palolo, waiomao valley PRESERVED_SPECIMEN tongg, r. collector number: s.n. Molokai BISH 11/29/1942
68 Dioscorea pentaphylla L. Kapakahi Gulch Koa zone in ope kukui forest. PRESERVED_SPECIMEN Egler, F.E. 37-215 Oahu BISH 7/22/1937
69 Dioscorea pentaphylla L. North Molokai; slopes of Waiahookalo Stream valley Steep, rocky slopes w/secondary forest of Psidium guajava, Syzygium malaccense & native remnants incl. Diospyros sandwicensis, Metrosideros & Nestegis w/understory of Kalanchoe pinnata & Zingiber zerumbet. Common along stream. PRESERVED_SPECIMEN Lorence, D.H. 5419 Molokai BISH 7/31/1987
70 Dioscorea pentaphylla L. manoa valley, waiakeakua, tubens PRESERVED_SPECIMEN wentworth, c.h. Oahu BISH 8/8/1935
71 Dioscorea pentaphylla L. Palolo Valley, Kaau Crater, frequent along trail on way to crater Koa zone. PRESERVED_SPECIMEN Egler, F.E. 37-309 Oahu BISH 7/29/1937
72 Dioscorea pentaphylla L. palolo valley, kaau crater, frequent along trail on way to crater Along ditch in rice field PRESERVED_SPECIMEN egler, f.e. collector number: 37-309 Oahu BISH 7/29/1937
73 Dioscorea pentaphylla L. koolau range; kawailoa, kawaiiki ditch trail, side gulch Rain forest. PRESERVED_SPECIMEN wilson, k.a. collector number: 12 Oahu BISH 10/14/1951
74 Dioscorea pentaphylla L. Koolau Mts.; Koolauloa, Hauula, "Hidden Valley", above Swanzy Park.tch Trail, side gulch Vine gowing on Pisonia PRESERVED_SPECIMEN Teraoka, W. 360 Oahu BISH 10/9/1980
75 Dioscorea pentaphylla L. Lihue-Koloa Forest Reserve; 3.2 miles NW of Wailua Experiment Station Vine climbing on Aleurites & Hibiscus overstory. In shade w/Zingiber, Nephrolepis, Oplismenus. PRESERVED_SPECIMEN Ishikawa, S. 277 Kauai BISH 8/20/1973
76 Dioscorea pentaphylla L. Punaluu; Castle Trail Loose rocky soil, steep slope, ohia woods. PRESERVED_SPECIMEN St. John, H. 23376 Oahu BISH 5/23/1948
77 Dioscorea pentaphylla L. west niu gulch, at branch in stream Koa zone in opening in forest. PRESERVED_SPECIMEN egler, f.e. collector number: 37-195 Oahu BISH 7/20/1937
78 Dioscorea pentaphylla L. Palolo Valley PRESERVED_SPECIMEN Forbes, C.N. 1968.O Oahu BISH 10/22/1914
79 Dioscorea pentaphylla L. West Niu Gulch, at branch in stream Koa zone in opening in forest. PRESERVED_SPECIMEN Egler, F.E. 37-195 Oahu BISH 7/20/1937
80 Dioscorea pentaphylla L. wailau valley floor PRESERVED_SPECIMEN higashino, p.k. allen, m.s. collector number: 10374 Molokai BISH 8/13/1984
81 Dioscorea pentaphylla L. east maui, hana district, kalahu point, ptbg kahanu garden In remnant Pandanus forest SSE of Pi`ilanihale Heiau in rough a`a lava. Common throughout forest. PRESERVED_SPECIMEN flynn, t. collector number: 2487 Maui BISH 10/20/1987
82 Dioscorea pentaphylla L. Halepuaa Forest Reserve in the Puna District PRESERVED_SPECIMEN Clarke, G. 79.205 Hawaii BISH 3/7/1979
83 Dioscorea pentaphylla L. wainiha valley, near dam wet; Metrosideros, Psychotria PRESERVED_SPECIMEN stone, b.c. collector number: 1505 Molokai BISH 12/26/1956
84 Dioscorea pentaphylla L. Ewa Dist.; Waimalu Metrosideros forest. PRESERVED_SPECIMEN Grant, M.L. 7338 Oahu BISH 9/9/1934
85 Dioscorea pentaphylla L. halepuaa forest reserve in the puna district PRESERVED_SPECIMEN clarke, g. collector number: 79.205 Hawaii BISH 3/7/1979
86 Dioscorea pentaphylla L. haupu, kipu Moist forest. Climbing over trees. PRESERVED_SPECIMEN st.john, h. fosberg, f.r. collector number: 13672 Kauai BISH 12/25/1933
87 Dioscorea pentaphylla L. 2.5km w of hana hwy & waikaloa rd, in the kawaipapa area Tropical lowland forest, planted with introduced trees PRESERVED_SPECIMEN c. r. annable 3237 NY 1/15/1997
88 Dioscorea pentaphylla L. 2.5km W of Hana Hwy & Waikaloa Rd, in the Kawaipapa area Tropical lowland forest, planted with introduced trees PRESERVED_SPECIMEN C. R. Annable 3237 NY 1/15/1997
89 Dioscorea pentaphylla L. Kalahu Point. PTBG Kahanu Garden. SSE of Pi'ilanihale Heiau on rough aa lava. Elev. 0-50 ft. In remnant Pandanus forest. PRESERVED_SPECIMEN Tim Flynn 2487 Maui PTBG 10/20/1987
90 Dioscorea pentaphylla L. Kealia Forest Reserve; Makaleha Ditch intake on Makaleha Stream. PRESERVED_SPECIMEN Tim Flynn 1332 Kauai PTBG 10/25/1985
91 Dioscorea pentaphylla L. West Maui, Honolua Valley. Lowland Wet Riparian Forest. PRESERVED_SPECIMEN Hank Oppenheimer H100604 Maui PTBG 10/4/2006
92 Dioscorea pentaphylla L. Kuilau Trail, below Keahua Arboretum. Disturbed lowland windward Forest PRESERVED_SPECIMEN Natalia Tangalin 3559 Kauai PTBG 11/22/2013
93 Dioscorea pentaphylla L. Waimalu, Koolau Range, at middle elevation PRESERVED_SPECIMEN Degener, O.; Park; Potter; Shigeura; Takamoto, -. 10110 Oahu US 11/3/1935
94 Dioscorea pentaphylla L. Slopes of Waiohookalo Stream Valley (N. Molokai); steep, rocky slopes. Alt. 100-180 m. With secondary forest. PRESERVED_SPECIMEN David H. Lorence 5419 Molokai PTBG 7/31/1987
95 Dioscorea pentaphylla L. waimalu, koolay range, oahu, territory of hawaii PRESERVED_SPECIMEN o. degener 10110 Hawaii US 11/3/1935
96 Dioscorea pentaphylla L. coast nw of hilo PRESERVED_SPECIMEN wilkes explor. exped. s.n. Hawaii US
97 Dioscorea pentaphylla L. honaunau PRESERVED_SPECIMEN a. s. hitchcock 14545 Hawaii US 9/3/1916
98 Dioscorea pentaphylla L. waimalu, koolau range, at middle elevation PRESERVED_SPECIMEN o. degener, park, potter, shigeura & -. takamoto 10110 Oahu US 11/3/1935
99 Dioscorea pentaphylla L. 3.2 mi NW of Wailua Experiment Station, Lihue-Koloa Forest Reserve PRESERVED_SPECIMEN Ishikawa, S. S. 277 Kauai US 8/20/1973
100 Dioscorea pentaphylla L. Honaunau PRESERVED_SPECIMEN Hitchcock, A. S. 14545 Hawaii US 9/3/2016
101 Dioscorea pentaphylla L. oahu PRESERVED_SPECIMEN st.john, h. collector number: 24834 Oahu BISH
102 Dioscorea pentaphylla L. PRESERVED_SPECIMEN degener, otto 10110 USCH 11/3/1935
103 Dioscorea pentaphylla L. Wailau Valley Climbing over shrubs. PRESERVED_SPECIMEN Rock, J.F.C. 70221 Molokai BISH 4/20/1910
104 Dioscorea pentaphylla L. waioli valley PRESERVED_SPECIMEN forbes, c.n. collector number: 91.k Kauai BISH 7/29/1909
105 Dioscorea pentaphylla L. East Maui, Hana District, Kalahu Point, PTBG Kahanu Garden In remnant Pandanus forest SSE of Pi`ilanihale Heiau in rough a`a lava. Common throughout forest. PRESERVED_SPECIMEN Flynn, T. 2487 Maui BISH 10/20/1987
106 Dioscorea pentaphylla L. koolau range; waimalu Flowering only bare landslide at middle elevation. PRESERVED_SPECIMEN degener, o. park, k.k.; potter; shigeura; takamoto collector number: 10110 Oahu BISH 11/3/1935
107 Dioscorea pentaphylla L. palolo valley PRESERVED_SPECIMEN forbes, c.n. collector number: 1968.o Oahu BISH 10/22/1914
108 Dioscorea pentaphylla L. wainiha; wainiha valley, bottom of small side gulch Moist lower forest. Climbing over trees. PRESERVED_SPECIMEN st.john, h. collector number: 13908 Kauai BISH 1/1/1934
109 Dioscorea pentaphylla L. head of hanakoa valley Wet woods. PRESERVED_SPECIMEN st.john, h. collector number: 24968 Kauai BISH 1/3/1953
110 Dioscorea pentaphylla L. 3.2 mi nw of wailua experiment station, lihue-koloa forest reserve PRESERVED_SPECIMEN s. s. ishikawa 277 Kauai US 8/20/1973
111 Dioscorea pentaphylla L. Wailau Valley floor PRESERVED_SPECIMEN Higashino, P.K. 10374 Molokai BISH 8/13/1984
112 Dioscorea pentaphylla L. Waimano Trail PRESERVED_SPECIMEN Kerr, M. 439-1 Oahu BISH 10/7/1945
113 Dioscorea pentaphylla L. Koolau Range; Palolo, Waiomao Valley Climbing over low shrubs, dense undergrowth. PRESERVED_SPECIMEN Kuykendall, J.R. 45 Oahu BISH 11/29/1942
114 Dioscorea pentaphylla L. Gulch west of Ualapue Rocky woods. PRESERVED_SPECIMEN Degener, O. 5562 Molokai BISH 6/30/1928
115 Dioscorea pentaphylla L. Koolau Range; Palolo, Waiomao Valley Intermediate moist. PRESERVED_SPECIMEN Nishida, T. s.n. Oahu BISH 10/5/1943
116 Dioscorea pentaphylla L. punaluu; castle trail Loose rocky soil, steep slope, ohia woods. PRESERVED_SPECIMEN st.john, h. collector number: 23376 Oahu BISH 5/23/1948
117 Dioscorea pentaphylla L. ewa dist.; waimalu Metrosideros forest. PRESERVED_SPECIMEN grant, m.l. collector number: 7338 Oahu BISH 9/9/1934
118 Dioscorea pentaphylla L. east maui, hana district, 2.5 km w of hana hwy & waikoloa rd in the kawaipapa area Tropical lowland forest planted w. introduced trees. PRESERVED_SPECIMEN annable, c.r. hobdy, r. collector number: 3237 Maui BISH 1/15/1997
119 Dioscorea pentaphylla L. waimalu Damp shaded rocks in moderately dry mesic ohia forest w/ Aleurites and Eugenia malaccensis common in wetter gullies PRESERVED_SPECIMEN brown, f.b.h. collector number: 11282 Oahu BISH 12/31/1925
120 Dioscorea pentaphylla L. lihue-koloa forest reserve; 3.2 miles nw of wailua experiment station Vine climbing on Aleurites & Hibiscus overstory. In shade w/Zingiber, Nephrolepis, Oplismenus. PRESERVED_SPECIMEN ishikawa, s. collector number: 277 Kauai BISH 8/20/1973
121 Dioscorea pentaphylla L. waimano trail PRESERVED_SPECIMEN kerr, m. collector number: 439-1 Oahu BISH 10/7/1945
122 Dioscorea pentaphylla L. ahupuaa of Halepuaa Forest Reserve in the Puna District. Found growing on the side of the old railroad On the side of the old railroad. PRESERVED_SPECIMEN ESP Field Crew s.n. Hawaii BISH 8/15/1979
123 Dioscorea pentaphylla L. head of wailau valley Rain forest near banana patch. PRESERVED_SPECIMEN st.john, h. et al. collector number: 23531 Molokai BISH 12/27/1948
124 Dioscorea pentaphylla L. Palolo, Waiomao Valley Open shrubby area in valley. PRESERVED_SPECIMEN Tongg, R. s.n. Oahu BISH 11/29/1942
125 Dioscorea pentaphylla L. north molokai; slopes of waiahookalo stream valley Steep, rocky slopes w/secondary forest of Psidium guajava, Syzygium malaccense & native remnants incl. Diospyros sandwicensis, Metrosideros & Nestegis w/understory of Kalanchoe pinnata & Zingiber zerumbet. Common along stream. PRESERVED_SPECIMEN lorence, d.h. wagner, w.l.; perlman, s.; montgomery, s. collector number: 5419 Molokai BISH 7/31/1987
126 Dioscorea pentaphylla L. koolau range; palolo, waiomao valley Intermediate moist. PRESERVED_SPECIMEN nishida, t. collector number: s.n. Oahu BISH 10/5/1943
127 Dioscorea pentaphylla L. ahupuaa of halepuaa forest reserve in the puna district. found growing on the side of the old railroad On the side of the old railroad. PRESERVED_SPECIMEN esp field crew collector number: s.n. Hawaii BISH 8/15/1979
128 Dioscorea pentaphylla L. Wainiha; Wainiha Valley, bottom of small side gulch Moist lower forest. Climbing over trees. PRESERVED_SPECIMEN St. John, H. 13908 Kauai BISH 1/1/1934
129 Dioscorea pentaphylla L. Head of Wailau Valley Rain forest near banana patch. PRESERVED_SPECIMEN St. John, H. 23531 Molokai BISH 12/27/1948
130 Dioscorea pentaphylla L. kapakahi gulch Koa zone in ope kukui forest. PRESERVED_SPECIMEN egler, f.e. collector number: 37-215 Oahu BISH 7/22/1937
131 Dioscorea pentaphylla L. east maui, hana district, hana forest reserve, hana ranch lands, mauka of keakaamanu Alien forest slopes dominated by Psidium guajava and Ardisia elliptica. PRESERVED_SPECIMEN imada, c.t. legrande, m.; jansen, s.; crummer, j. collector number: 2002-80 Maui BISH 11/11/2002
132 Dioscorea pentaphylla L. Waioli Valley PRESERVED_SPECIMEN Forbes, C.N. 91.K Kauai BISH 7/29/1909
133 Dioscorea pentaphylla L. Waimalu PRESERVED_SPECIMEN Brown, F.B.H. 11282 Oahu BISH 12/31/1925
134 Dioscorea pentaphylla L. koolau range; palolo, waiomao valley Climbing over low shrubs, dense undergrowth. PRESERVED_SPECIMEN kuykendall, j.r. collector number: 45 Oahu BISH 11/29/1942
135 Dioscorea pentaphylla L. Koolau Range; Kawailoa, Kawaiiki Ditch Trail, side gulch Rain forest. PRESERVED_SPECIMEN Wilson, K.A. 12 Oahu BISH 10/14/1951
136 Dioscorea pentaphylla L. ahupuaa of Halepuaa Forest Reserve in the Puna District. Found growing on the side of the old railroad Tropical lowland forest, planted with introduced trees PRESERVED_SPECIMEN C. R. Annable 3237 Hawaii HAPI 1/15/1997
137 Dioscorea pentaphylla L. wailau valley Climbing over shrubs. PRESERVED_SPECIMEN rock, j.f.c. collector number: 70221 Molokai BISH 4/20/1910
138 Dioscorea pentaphylla L. gulch west of ualapue Rocky woods. PRESERVED_SPECIMEN degener, o. collector number: 5562 Molokai BISH 6/30/1928
139 Dioscorea pentaphylla L. East Maui, Hana District, 2.5 km W of Hana Hwy & Waikoloa Rd in the Kawaipapa area Tropical lowland forest planted w. introduced trees. PRESERVED_SPECIMEN Annable, C.R. 3237 Maui BISH 1/15/1997
140 Dioscorea pentaphylla L. East Maui, Hana District, Hana Forest Reserve, Hana Ranch lands, mauka of Keakaamanu Tropical lowland forest, planted with introduced trees UNKNOWN C. R. Annable Maui HAPI 1/15/1997
141 Dioscorea pentaphylla L. Koolau Range; Waimalu Flowering only bare landslide at middle elevation. PRESERVED_SPECIMEN Degener, O. 10110 Oahu BISH 11/3/1935
142 Dioscorea pentaphylla L. Head of Hanakoa Valley Wet woods. PRESERVED_SPECIMEN St. John, H. 24968 Kauai BISH 1/3/1953
143 Dioscorea pentaphylla L. koolau range, kawailoa, kawaiiki ditch trail Moist wooded side gulch, under Aleurites and Metrosideros. PRESERVED_SPECIMEN st.john, h. collector number: 24704 Oahu BISH 10/14/1951
144 Dioscorea pentaphylla L. East Maui, Hana District, 2.5 km W of Hana Hwy & Waikoloa Rd in the Kawaipapa area Tropical lowland forest planted w. introduced trees. PRESERVED_SPECIMEN Annable, C.R. Hobdy, R. Collector Number: 3237 Maui BISH 1/15/1997
145 Dioscorea pentaphylla L. East Maui, Hana District, Hana Forest Reserve, Hana Ranch lands, mauka of Keakaamanu Alien forest slopes dominated by Psidium guajava and Ardisia elliptica. PRESERVED_SPECIMEN Imada, C.T. 2002-80 Maui BISH 11/11/2002
146 Dioscorea pentaphylla L. Halepuaa Forest Reserve in the Puna District PRESERVED_SPECIMEN Clarke, G. Collector Number: 79.205 Hawaii BISH 3/7/1979
147 Dioscorea pentaphylla L. Koolau Range, Kawailoa, Kawaiiki Ditch trail Moist wooded side gulch, under Aleurites and Metrosideros. PRESERVED_SPECIMEN St. John, H. 24704 Oahu BISH 10/14/1951
148 Dioscorea pentaphylla L. East Maui, Hana District, Hana Forest Reserve, Hana Ranch lands, mauka of Keakaamanu Alien forest slopes dominated by Psidium guajava and Ardisia elliptica. PRESERVED_SPECIMEN Imada, C.T. LeGrande, M.; Jansen, S.; Crummer, J. Collector Number: 2002-80 Maui BISH 11/11/2002
149 Dioscorea pentaphylla L. Wainiha Valley, near dam PRESERVED_SPECIMEN Stone, B.C. 1505 Kauai BISH 12/26/1956
150 Dioscorea pentaphylla L. Kawailoa, Kawaiiki Ditch Trail PRESERVED_SPECIMEN St. John, H. 24834 Oahu BISH 7/13/1952
151 Dioscorea pentaphylla L. koolau mts.; koolauloa, hauula, "hidden valley", above swanzy park.tch trail, side gulch Vine gowing on Pisonia PRESERVED_SPECIMEN teraoka, w. nagata, k.; whiteaker, l.; silva, b. collector number: 360 Oahu BISH 10/9/1980
152 Dioscorea pentaphylla L. ahupuaa of Halepuaa Forest Reserve in the Puna District. Found growing on the side of the old railroad On the side of the old railroad. PRESERVED_SPECIMEN ESP Field Crew Collector Number: s.n. Hawaii BISH 8/15/1979
153 Dioscorea pentaphylla L. Manoa Valley, Waiakeakua, PRESERVED_SPECIMEN Wentworth, C.H. s.n. Oahu BISH 4/18/1905
154 Dioscorea pentaphylla L. Haupu, Kipu Moist forest. Climbing over trees. PRESERVED_SPECIMEN St. John, H. 13672 Kauai BISH 12/25/1933
155 Dioscorea pentaphylla L. palolo, waiomao valley PRESERVED_SPECIMEN tongg, r. collector number: s.n. Molokai BISH 11/29/1942
156 Dioscorea pentaphylla L. Kapakahi Gulch Koa zone in ope kukui forest. PRESERVED_SPECIMEN Egler, F.E. 37-215 Oahu BISH 7/22/1937
157 Dioscorea pentaphylla L. North Molokai; slopes of Waiahookalo Stream valley Steep, rocky slopes w/secondary forest of Psidium guajava, Syzygium malaccense & native remnants incl. Diospyros sandwicensis, Metrosideros & Nestegis w/understory of Kalanchoe pinnata & Zingiber zerumbet. Common along stream. PRESERVED_SPECIMEN Lorence, D.H. 5419 Molokai BISH 7/31/1987
158 Dioscorea pentaphylla L. manoa valley, waiakeakua, tubens PRESERVED_SPECIMEN wentworth, c.h. Oahu BISH 8/8/1935
159 Dioscorea pentaphylla L. Palolo Valley, Kaau Crater, frequent along trail on way to crater Koa zone. PRESERVED_SPECIMEN Egler, F.E. 37-309 Oahu BISH 7/29/1937
160 Dioscorea pentaphylla L. palolo valley, kaau crater, frequent along trail on way to crater Along ditch in rice field PRESERVED_SPECIMEN egler, f.e. collector number: 37-309 Oahu BISH 7/29/1937
161 Dioscorea pentaphylla L. koolau range; kawailoa, kawaiiki ditch trail, side gulch Rain forest. PRESERVED_SPECIMEN wilson, k.a. collector number: 12 Oahu BISH 10/14/1951
162 Dioscorea pentaphylla L. Koolau Mts.; Koolauloa, Hauula, "Hidden Valley", above Swanzy Park.tch Trail, side gulch Vine gowing on Pisonia PRESERVED_SPECIMEN Teraoka, W. 360 Oahu BISH 10/9/1980
163 Dioscorea pentaphylla L. Lihue-Koloa Forest Reserve; 3.2 miles NW of Wailua Experiment Station Vine climbing on Aleurites & Hibiscus overstory. In shade w/Zingiber, Nephrolepis, Oplismenus. PRESERVED_SPECIMEN Ishikawa, S. 277 Kauai BISH 8/20/1973
164 Dioscorea pentaphylla L. Punaluu; Castle Trail Loose rocky soil, steep slope, ohia woods. PRESERVED_SPECIMEN St. John, H. 23376 Oahu BISH 5/23/1948
165 Dioscorea pentaphylla L. west niu gulch, at branch in stream Koa zone in opening in forest. PRESERVED_SPECIMEN egler, f.e. collector number: 37-195 Oahu BISH 7/20/1937
166 Dioscorea pentaphylla L. Palolo Valley PRESERVED_SPECIMEN Forbes, C.N. 1968.O Oahu BISH 10/22/1914
167 Dioscorea pentaphylla L. West Niu Gulch, at branch in stream Koa zone in opening in forest. PRESERVED_SPECIMEN Egler, F.E. 37-195 Oahu BISH 7/20/1937
168 Dioscorea pentaphylla L. wailau valley floor PRESERVED_SPECIMEN higashino, p.k. allen, m.s. collector number: 10374 Molokai BISH 8/13/1984
169 Dioscorea pentaphylla L. east maui, hana district, kalahu point, ptbg kahanu garden In remnant Pandanus forest SSE of Pi`ilanihale Heiau in rough a`a lava. Common throughout forest. PRESERVED_SPECIMEN flynn, t. collector number: 2487 Maui BISH 10/20/1987
170 Dioscorea pentaphylla L. Halepuaa Forest Reserve in the Puna District PRESERVED_SPECIMEN Clarke, G. 79.205 Hawaii BISH 3/7/1979
171 Dioscorea pentaphylla L. wainiha valley, near dam wet; Metrosideros, Psychotria PRESERVED_SPECIMEN stone, b.c. collector number: 1505 Molokai BISH 12/26/1956
172 Dioscorea pentaphylla L. Ewa Dist.; Waimalu Metrosideros forest. PRESERVED_SPECIMEN Grant, M.L. 7338 Oahu BISH 9/9/1934
173 Dioscorea pentaphylla L. halepuaa forest reserve in the puna district PRESERVED_SPECIMEN clarke, g. collector number: 79.205 Hawaii BISH 3/7/1979
174 Dioscorea pentaphylla L. haupu, kipu Moist forest. Climbing over trees. PRESERVED_SPECIMEN st.john, h. fosberg, f.r. collector number: 13672 Kauai BISH 12/25/1933
175 Dioscorea pentaphylla L. 2.5km w of hana hwy & waikaloa rd, in the kawaipapa area Tropical lowland forest, planted with introduced trees PRESERVED_SPECIMEN c. r. annable 3237 NY 1/15/1997
176 Dioscorea pentaphylla L. 2.5km W of Hana Hwy & Waikaloa Rd, in the Kawaipapa area Tropical lowland forest, planted with introduced trees PRESERVED_SPECIMEN C. R. Annable 3237 NY 1/15/1997
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